Lycaena

Lycaena Fabricius, 1807

Lycaena Fabricius, 1807; Mag. f. Insektenk. 6: 285; TS: Papilio phlaeas Linnaeus
Apangea Zhdanko, 1995; Ent. Obozr. 74 (3): 658; TS: Chrysophanus pang Oberthür
Lafron Grishin, 2020 (Lycaeninae); Taxonomic Rep. 8 (7): 12; TS: Papilio orus Stoll

Apangea; [NHM-BMW]
Apangea (Lycaeninae); Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 12
Lafron; [iNaturalist]
Lycaena (Lycaena); [P&N23]
Lycaena; Godman & Salvin, [1887], Biol. centr.-amer., Lep. Rhop. 2: 102; [P&N23]; [CoL] ; [iNaturalist] ; [iNaturalist] ; [NHM-BMW] ; [NHM-BMW]
Lycaena (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena (Lycaenidae); Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 263; [BRU2], 123
Lycaena (Lycaeninae); [BK], 212; [NL4A], 137; [AfrL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; [RFEL]; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Lycaena (Lycaenini); Pelham, 2008, J. Res. Lepid. 40: 188; Krupitsky, 2017, Zootaxa 4306 (1): 138 (list)

Subgenus Lycaena

Lycia Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6): 81 (repl. Lycaena Fabricius, 1807); TS: Papilio phlaeas Linnaeus
- = Lycaena; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; [NL4A], 137; [AfrL]
- = Subgenus Lycaena (Lycaena); Pelham, 2008, J. Res. Lepid. 40: 188
Migonitis Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6): 82 (repl. Lycaena Fabricius, 1807, preocc. Migonitis Rafinesque, 1815, Migonitis Hübner, 1816); TS: Papilio phlaeas Linnaeus
- = Lycaena; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [NL4A], 137; [AfrL]
- = Subgenus Lycaena (Lycaena); Pelham, 2008, J. Res. Lepid. 40: 188
Rumicia Tutt, 1906; Ent. Rec. J. Var. 18 (5): 131; TS: Papilio phlaeas Linnaeus
- = Lycaena; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [NL4A], 138; [AfrL]
- = Subgenus Lycaena (Lycaena); Pelham, 2008, J. Res. Lepid. 40: 188
Helleia Verity, 1943; Le Farfalle diurn. d'Italia 2: 20, 48; TS: Papilio helle Denis & Schiffermüller
- = Lycaena; [NL4A], 138

Subgenus Lycaena (Lycaena); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Pelham, 2008, J. Res. Lepid. 40: 188; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Helleia; [CoL] ; [NHM-BMW]
Helleia (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Helleia (Lycaeninae); Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Rumicia; [NHM-BMW]

Lycaena phlaeas (Linnaeus, 1761)

EU, NAF, AM, Temperate Asia, Japan, Abyssinia. See [maps] Gyllenvinge Small Copper Le Bronzé American Copper Liten guldvinge Czerwończyk żarek Pikkukultasiipi Common Copper Kleiner Feuerfalter Manto bicolor

Papilio phlaeas Linnaeus, 1761; Fauna Suecica (Edn 2): 285; TL: C. Sweden
Papilio virgaureae Scopoli, 1763; Ent. Carniolica: 180
Polyommatus melanophlaeas Guenée & Villiers, 1835; Tabl. Synop. 1: 36
Lycaena phlaeas vernus Zeller, 1847; Isis von Oken 1847 (2): 158
Lycaena phlaeas aestivus Zeller, 1847; Isis von Oken 1847 (2): 158
Polyommatus phlaeas var. schmidtii Gerhard, 1850; Versuch Mon. europ. Schmett. (3): 7, pl. 10, f. 3a-b
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 416
Polyommatus phlaeas var. cuprinus Peyerimhoff, 1862; Cat. Lep. Alsace: 13
Polyommatus phlaeas ab. albicans Fuchs, 1884; Stettin ent. Ztg. 45 (7-9): 249
Polyommatus phlaeas var. semi-schmidtii Robson, 1888; Young Naturalist 9: 180
Polyommatus phlaeas var.[?] fuscae Robson, 1888; Young Naturalist 9: 181
Polyommatus phlaeas var. posti-striatae Robson, 1888; Young Naturalist 9: 181
caeruleopunctata Staudinger, 1895²; Palaearkt. Grossschm. 1: 2
Polyommatus phlaeas ♀ ab. supra-minus-punctata Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 72-73
Polyommatus phlaeas ♀ ab. supra-radiata Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 74
Chrysophanus phlaeas fasciatus Cockerell, 1889; Entomologist 22 (311): 99
Polyommatus phlaeas ♂ ab. punctis nigris remotis Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 75
Polyommatus phlaeas ♂ ab. absque vitta marginali rubro-aurea Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 77
Lycaena phlaeas ab. intermedia Tutt, 1896; Brit. Butts: 153
Lycaena phlaeas ab. suffusa Tutt, 1896; Brit. Butts: 153
Lycaena phlaeas ab. obsoleta Tutt, 1896; Brit. Butts: 153
Lycaena phlaeas ab. radiata Tutt, 1896; Brit. Butts: 153
Polyommatus phlaeas ab. exticta Fuchs, 1899; Jb. Nassau Ver. Nat. 52: 121
Polyommatus phlaeas var. transiens Fuchs, 1899; Jb. Nassau Ver. Nat. 52: 121
Chrysophanus phlaeas ab. kochi Strand, 1902; Nyt Mag. Naturv. 40: 162
Chrysophanus phlaeas ab. parvipuncta Strand, 1902; Nyt Mag. Naturv. 40: 163
Chrysophanus phlaeas f. coeruleopuncta Strand, 1902; Nyt Mag. Naturv. 40: 163
Chrysophanus phlaeas f. angustimargo Courvoisier, 1903; Mitt. schweiz. ent. Ges. 11: 25
Chrysophanus phlaeas ab. spoliata Schultz, 1903; Nyt mag. naturv. 41: 25
Chrysophanus phlaeas var. hübneri Oberthür, 1905; Bull. Soc. ent. Fr. 1905: 56
Chrysophanus phlaeas ab. auronitens Schultz, 1905; Ent. Zs. 18 (34): 134
Chrysophanus phlaeas v. eleus ab. cuneifera Schultz, 1905; Ent. Zs. 18 (34): 134
Rumicia phlaeas ab. minor Tutt, 1906; Nat. Hist. Br. Lepid. 8: 353
Rumicia phlaeas ab. ignita Tutt, 1906; Nat. Hist. Br. Lepid. 8: 353
Rumicia phlaeas ab. major Tutt, 1906; Nat. Hist. Br. Lepid. 8: 353
Rumicia phlaeas ab. hübneri Tutt, 1906; Nat. Hist. Br. Lepid. 8: 355
Rumicia phlaeas ab. webbi Tutt, 1906; Nat. Hist. Br. Lepid. 8: 355
Rumicia phlaeas ab. infuscata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 355
Rumicia phlaeas ab. alba Tutt, 1906; Nat. Hist. Br. Lepid. 8: 355
Rumicia phlaeas ab. dextroalba Tutt, 1906; Nat. Hist. Br. Lepid. 8: 357
Rumicia phlaeas ab. anteroalba Tutt, 1906; Nat. Hist. Br. Lepid. 8: 357
Rumicia phlaeas ab. posteroalba Tutt, 1906; Nat. Hist. Br. Lepid. 8: 357
Rumicia phlaeas ab. alba-caudata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 357
Rumicia phlaeas ab. schmidtii-caudata [= schmidtiicaudata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 358
Rumicia phlaeas ab. intermedia-caudata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 359
Rumicia phlaeas ab. typica-caudata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 359
Rumicia phlaeas ab. ignita-caudata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 360
Rumicia phlaeas ab. purpureotincta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 360
Rumicia phlaeas ab. bipunctata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 360
Rumicia phlaeas ab. unipunctata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 361
Rumicia phlaeas ab. impunctata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 361
Rumicia phlaeas ab. remota Tutt, 1906; Nat. Hist. Br. Lepid. 8: 361
Rumicia phlaeas ab. remota-juncta [= remotajuncta] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 361
Rumicia phlaeas ab. magnipuncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 361
Rumicia phlaeas ab. juncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 362
Rumicia phlaeas ab. juncta-caudata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 362
Rumicia phlaeas ab. extensa Tutt, 1906; Nat. Hist. Br. Lepid. 8: 363
Rumicia phlaeas ab. extensa-conjuncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 363
Rumicia phlaeas ab. centriconjuncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 364
Rumicia phlaeas ab. latomarginata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 366
Rumicia phlaeas ab. nigroapicata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 366
Rumicia phlaeas ab. basilipuncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 366
Rumicia phlaeas ab. cupreopunctata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas ab. subradiata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas ab. fasciata-caeruleopunctata [= fasciatacaeruleopunctata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas ab. eleus-caeruleopunctata [= eleuscaeruleopunctata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas var. timeus-caeruleopunctata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas ab. obsoleta-caeruleopunctata [= obsoletacaeruleopunctata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 368
Rumicia phlaeas ab. initia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 374
Rumicia phlaeas ab. initia-caudata [= initiacaudata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 374
Rumicia phlaeas ab. fuscata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 378
Rumicia phlaeas ab. fuscata-caudata [= fuscacaudata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 378
Rumicia phlaeas ab. infra-extensa [= infraextensa] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 379
Rumicia phlaeas ab. disco-juncta [= discojuncta] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 380
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 142
Rumicia phlaeas ab. infra-radiata [= infraradiata] Tutt, 1906; Nat. Hist. Br. Lepid. 8: 380
Lycaena phlaeas ab. parallela Courvoisier, 1907; Zs. wiss. Insektenbiol. 3 (2): 36
Chrysophanus phlaeas ab. caeca Courvoisier, 1907; Zs. wiss. Insektenbiol. 3 (3): 75
Rumicia phlaeas ab. addenda Williams, 1911; Ent. Rec. J. Var. 23: 275
ab. basijuncta Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 48
ab. paucipuncta Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 63
Chrysophanus phlaeas ab. semialba Strand, 1912; Ent. Zs. 25 (50): 257
radiata Frohawk, 1914²; Brit. Butts.: pl. 46, f. 29
Chrysophanus phlaeas f. paucimaculata Vorbrodt, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 444
Rumicia phlaeas r. varieleus Verity, 1920; Ent. Rec. J. Var. 32: 5
Rumicia phlaeas r. nigrioreleus Verity, 1920; Ent. Rec. J. Var. 32: 5
Chrysophanus phlaeas f. rubrohastata Stauder, 1923; Zs. Wiss. InsekBiol. 18 (3/4): 68
hyperborea (Ford, 1924) (Heodes); Trans. ent. Soc.: 739
minor (Derenne, 1926) (Chrysophanus); Lambillionea 26: 4
coreuleabasalis (Andres, 1931) (Chrysophanus); Bull. Soc. ent. Egypte 1930[?]: 212
berviniensis (Smith, 1932) (Heodes); Entomologist 65: 241
couteaui (Pionneau, 1938) (Chrysophanus); Misc. ent. 39: 46
semiobscurata Frohawk, 1938; Var. Brit. Butterfl. : pl. 34, f. 1-2
auroradiata Bright & Leeds, 1938; Monogr. Br. Ab. Chalk-hill Blue: 117
partimauroradiata Bright & Leeds, 1938; Monogr. Br. Ab. Chalk-hill Blue: 117
rufomargo Leeds, 1941; Monogr. Br. Ab. Chalk-hill Blue: 142
antibasijuncta Leeds, 1941; Monogr. Br. Ab. Chalk-hill Blue: 143
apicalisjuncta Leeds, 1941; Monogr. Br. Ab. Chalk-hill Blue: 143
ultrarufomargo Leeds, 1941; Monogr. Br. Ab. Chalk-hill Blue: 143
Lycaena phlaeas f. infrapallida Verity, 1943; Le Farfalle diurn. d'Italia 2: 52; TL: Italy
Lycaena phlaeas f. aestivaecaudata Verity, 1943; Le Farfalle diurn. d'Italia 2: 55; TL: Italy
Chrysophanus phlaeas ab. major Schnaider, 1950; Polskie Pismo. Ent. 19 (3-4): 252
Lycaena phlaeas f. rectaserie Lempke, 1954; Tijdschr. Ent. 97: 331
Lycaena phlaeas f. infrarufolineata Lempke, 1954; Tijdschr. Ent. 97: 331
basilijuncta Beuret, 1954; Lycaeniden Schweiz 1: 68
caeruleofasciata Holmes, 1978; Ent. Gaz. 29: 78

Lycaena (Lycaena) phlaeas; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Pelham, 2008, J. Res. Lepid. 40: 188; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; [RFEL]
Lycaena juncta; [CoL]
Lycaena melanophlaeas; [CoL]
Lycaena nigrioreleus; [CoL]
Lycaena phlaeas aestivaecaudata; [CoL]
Lycaena phlaeas basilijuncta; [CoL]
Lycaena phlaeas coeruleabasalis; [CoL]
Lycaena phlaeas hyperborea; [CoL]
Lycaena phlaeas infrapallida; [CoL]
Lycaena phlaeas infrarfolineata; [CoL]
Lycaena phlaeas rectaserie; [CoL]
Lycaena phlaeas schmidtii; [CoL]
Lycaena phlaeas vernus; [CoL]
Lycaena phlaeas; [BIR], 301; [SPRK]; [H&R]; [BARU], #343; [NACL], #4251; [NAB]; [EBW]; [BOW]: pl. 9, f. 39, 40; [MRS], 663; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BRU2]: 123, pl. 57, f. 7-39; [Opler]; [Otakar Kudrna]; Yakovlev, 2012, Nota lepid. 35 (1): 67; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [BAMONA] ; [SVF] ; [GALEPS] ; [Mario Maier]
Lycaena posteroalba; [CoL]
Lycaena suffusa; [CoL]
Lycaena vernus; [CoL]
Polyommatus phlaeas melanophlaeas; Oberthür, 1896, Étud. d'Ent. 20: pl. 5, f. 76
Polyommatus phlaeas schmidtii; Oberthür, 1896, Étud. d'Ent. 20: pl. 5, f. 70-71
Polyommatus phlaeas; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 365

646x554 (~56Kb) SPAIN: Teneriffa, 1-5.4.1996, Photo © Valentina Anderson	512x425 (~32Kb) SPAIN: Teneriffa, 1-5.4.1996, Photo © Valentina Anderson	500x450 (~58Kb) West Down, Devon, UK 12/06/92, Photo © Simon Coombes
450x381 (~26Kb) upperside France, Bretagne (Brittany), Bois De Lorge, 9 July 1997, Photo © Simon Coombes	500x439 (~26Kb) upperside Finland, Tampere, Rusko, 6820:339, 11.8.1996, Photo © Tero Piirainen	500x581 (~50Kb) underside Finland, Kangasala, Savo, 6815:338, 18.8.1996, Photo © Tero Piirainen
512x768 (~54Kb) upperside Germany, Baden-Wuerttemberg, Oetisheim, 8° 49' E, 48° 58' N, altitude 300m, 26.07.1998 (sucking on Origanum vulgare), Photo © Karl Hofsäß	512x768 (~47Kb) underside Germany, Baden-Wuerttemberg, Oetisheim, 8° 49' E, 48° 58' N, altitude 300m, 26.07.1998 (sucking on Origanum vulgare), Photo © Karl Hofsäß	identification uncertain 800x708 (~111Kb) upperside Ukraine, Ivano Frankovsk, near Kalush, Stankova, 24.10.2001, Photo © Erich Mangl
435x290 (~23Kb) Netherlands, Middenduin, Overdeen, May 2003, Photo © Peter Groenendijk	444x518 (~27Kb) A floweer clump, Chihaya-akasaka-mura, Minami-kawachi-gun, Osaka Prefecture, Japan. 3th August 2002, Photo © Oleg Kosterin	807x1170 (~181Kb) upperside Russian Federation, Krasnodarskii Krai, Adler, bank of Nizhneimeretinskaya harbour, 15 July 2008, Photo © Oleg Kosterin
560x520 (~83Kb) ♀ Russia, Moscow area, 27.8.2011 (36°25'E, 56°00'N), Photo © D. Smirnov	460x420 (~95Kb) ♂ Russia, Moscow area, 27.8.2011 (36°25'E, 56°00'N), Photo © D. Smirnov	450x298 (~21Kb)

L. p. phlaeas EU, W.Siberia, Caucasus, Transcaucasia

Lycaena phlaeas f. caerulopunctata Rühl, 1895; Palaearkt. Grossschm. 1: 218; TL: C.Asia
Chrysophanus phlaeas ab. oberthuri Blachier, 1908; Ann. Soc. ent. Fr. 77 (2): 217, pl. 4, f. 7; TL: (Morocco)

Lycaena phlaeas phlaeas; [BMAT]: 29, pl. 10, f. 1-11; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; [RFEL]; [iNaturalist]

600x435 (~34Kb) upperside Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

600x435 (~30Kb) underside Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

f. caeruleopunctata
448x396 (~28Kb) UK, W. SUFFOLK, Icklingham, King's Forest, GRID TL 7945 7330, 23.5.2004, Photo © Jonathan P. Tyler

L. p. timeus (Cramer, [1777])

Papilio timeus Cramer, [1777]; Uitl. Kapellen 2 (9-16): pl. 186, f. E, F
Hesperia eleus Fabricius, 1798; Ent. Syst. (Suppl.): 430
Polyommatus turcicus Gerhard, 1850; Versuch Mon. europ. Schmett. (2): 5, pl. 5, f. 5a-c

Lycaena phlaeas eleus; [iNaturalist]
Lycaena phlaeas timeus; [CoL] ; [iNaturalist]
Lycaena turcicus; [CoL]

L. p. hypophlaeas (Boisduval, 1852) Altai, S.Siberia, Amur, N.Ussuri, Massachusetts, New York, New Jersey

Polyommatus hypophlaeas Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 291; TL: "Nord de la California"
Chrysophanus americana D'Urban, 1860; Can. Nat. & Geol. 5 (4): 246 (Harris) (nom. nud.?)
Lycaena americana Harris, 1862; Treatise Ins. injur. Veget. (3rd edn): 273; TL: Massachusetts
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Chrysophanus phlaeas var. americana ab. ♀ fasciata Strecker, 1878; Synon. Cat. Macroplep.: 101; TL: "Florida" [?error]
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Chrysophanus americanus ab. fulliolus Hulst, 1886; Ent. Amer. 2 (9): 182; TL: [New York]
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Lycaena bacchus Scudder, 1889; Butts eastern U.S. Canada 2: 998 (Harris MS, nom. nud.)
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Heodes hypophlaeas ab. obliterata Scudder, 1889; Butts eastern U.S. Canada 2: 1001
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Chrysophanus phlaeas adrienne Maynard, 1891; Man. N. Amer. Butt., app.: 153; TL: Eastern Massachusetts
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
pusilla (Ksienschopolski, 1911) (Chrysophanus); Trudy obshch.Izsl. Volyni Zitomir 8: 56, pl. 1, f. 10
Heodes hypophlaeas f. caeca Reiff, 1913; Ent. News 24 (7): 306, pl. 10, f. 3-4; TL: West Roxbury, Mass.
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Chrysophanus hypohplaeas ab. octomaculata Dean, 1918; Lepidoptera 2 (8): 60; TL: [Missouri]
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Heodes hypophlaeas hypophlaeas ab. banksi Watson & Comstock, 1920; Bull. Amer. Mus. 42 (10): 454; TL: Lexington, Massachusetts
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Heodes hypophlaeas ab. fulvus Rummel, 1929; Bull. Brooklyn Ent. Soc. 23 (5): 268; TL: [New Jersey]
- = ; Pelham, 2008, J. Res. Lepid. 40: 189
Heodes hypophlaeas ab. neui Rummel, 1929; Bull. Brooklyn Ent. Soc. 23 (5): 268; TL: [New Jersey]
- = ; Pelham, 2008, J. Res. Lepid. 40: 189

Heodes hypophleas[sic, hypophlaeas]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 41
Lycaena (Lycaena) hypophlaeas hypophlaeas; [P&N23]
Lycaena (Lycaena) hypophlaeas; [P&N23]
Lycaena (Lycaena) phlaeas hypophlaeas; Pelham, 2008, J. Res. Lepid. 40: 188
Lycaena fasciata; [CoL]
Lycaena fulvus; [CoL]
Lycaena hypophlaeas caeca; [CoL]
Lycaena hypophlaeas hypophlaeas; [iNaturalist]
Lycaena hypophlaeas; [MPG] ; [CoL] ; [iNaturalist] ; [BOA]
Lycaena neui; [CoL]
Lycaena phlaeas americana; [NACL], #4251a; [CoL]
Lycaena phlaeas hypophlaeas; [NACL], #4251b; Ballmer & Pratt, 1989, J. Lep. Soc. 43 (1): 59; [BARU](note) , ; [BRU2], 124
obsoleta; Barnes & McDunnough, 1917, (missp.)

431x324 (~21Kb) ♀ Chelsea, Michigan, USA, May 2004, Photo © Christopher A. Rickards

L. p. chinensis (Felder, 1862) S.Ussuri, China (Shaanxi, Hupei, NE.China)

Chrysophanus phlaeas var. chinensis Felder, 1862; Verh. zool.-bot. Ges. Wien 12 (1/2): 488
matsumuranus (Bryk, 1946) (Heodes); Ark. Zool. 38 A (3): 55; TL: Korea

Lycaena phlaeas chinensis; [BARU](note) , ; [MRS], 663; [BRU2], 124; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]; Yoshino, 2019, Butterflies 80: 18 (note); [CoL] ; [iNaturalist]
Lycaena phlaeas matsumuranus; [CoL]

L. p. pseudophlaeas (Lucas, 1866) Ethiopia

Chrysophanus phlaeas pseudophlaeas Lucas, 1866; Ann. Soc. Ent. Fr. (4) 5: 499; TL: Abyssinia
Chrysophanus phlaeas var. menelicki Thierry-Mieg, 1910; Le Naturaliste 32 (561): 167; TL: Harrar, Abyssinia
- = ; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 264; [AfrL]

Lycaena phlaeas menelieki; [CoL]
Lycaena phlaeas pseudophlaeas; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 264; [BAFR], 525; [MRS], 663; de Jong & van Dorp, 2006, Ent. Ber. 66 (4): 125 (note,map); [AfrL]; [CoL] ; [iNaturalist]
Lycanea phlaeas pseudophlaeas; [BK], 213 (note)

L. p. feildeni (McLachlan, 1878) Ellesmere Is.

Chrysophanus phloeas[sic] feildeni McLachlan, 1878; J. Linn. Soc. Zool. Lond. 14: 111; TL: lat 81°41'N; Grinnell Land [SE.Ellesmere Is.]

Heodes hypophleas fieldeni[sic, feildeni]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 41
Lycaena (Lycaena) hypophlaeas feildeni; [P&N23]
Lycaena (Lycaena) phlaeas feildeni; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena feildeni; [CoL]
Lycaena hypophlaeas feildeni; [iNaturalist]
Lycaena phlaeas feildeni; [NACL], #4251e
Lycaena phlaeas fieldeni; [CoL]

L. p. stygianus (Butler, 1880) W.Pamirs

Chrysophanus stygianus Butler, 1880; Proc. zool. Soc. Lond. 1880 (3): 408, pl. 39, f. 6; TL: Candahar

Lycaena phlaeas stygiana; [BRU2], 123
Lycaena phlaeas stygianus; Bálint, 1992, Linn. Belg. 13 (8): 399; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]

L. p. baralacha (Moore, 1884) Ladak

Chrysophanus baralacha Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 25; TL: Baralacha Pass (16060ft), Ladak

Lycaena baralacha; [CoL]
Lycaena phlaeas baralacha; [iNaturalist]

L. p. oxiana (Grum-Grshimailo, 1890) Kopet-Dagh, Alai, Ghissar-Darvaz, Tian-Shan

Polyommatus phlaeas var. oxiana Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4: 365
turanica (Rühl, 1895) (Polyommatus); Palaearkt. Grossschm. 1: 747; TL: Tura
Lycaena phlaeas f. naruena Courvoisier, 1911; Ent. Zs. 24 (37): 202, (49):262
coccineus (Ford, 1924) (Heodes); Trans. ent. Soc.: 726

Lycaena phlaeas narua; [CoL]
Lycaena phlaeas naruena; [CoL]
Lycaena phlaeas oxiana; [BRU2], 123; [Oleg Kosterin]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [CoL] ; [iNaturalist]
Lycaena phlaeas turanica; [CoL]

533x400 (~65Kb) ♂ upperside A bank of the Shirkent River in its lower flow several km upstream of the kishlak Shirkent, the southern principal slope of the Ghissar mountain range, the Pamiro-Alai Mts., Tursun-Zade Province, Tadjikistan. 7th May 1989, Photo © Oleg Kosterin

L. p. comedarum (Grum-Grshimailo, 1890) E.Pamirs

Polyommatus phlaeas var. comedarum Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4: 366

Lycaena phlaeas comedarum; [BRU2], 123; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [CoL] ; [iNaturalist]

L. p. abbottii (Holland, 1892) Kenya, Tanzania, Malawi, Zambia

Chrysophanus abbottii Holland, 1892; Entomologist 25 (Suppl.): 90
Chrysophanus abbottii Holland, 1896; Proc. U.S. Nat. Mus. 18 (1062): 240, pl. 7, f. 1

Lycaena (Lycaena) abottii[sic, abbottii]; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena abbottii; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 264; [CoL]
Lycaena abotti[sic, abbottii]; [EBW]
Lycaena phlaeas abbotti; [BAFR], 525; [BK]: 213, pl. 21, f. 274
Lycaena phlaeas abbottii; de Jong & van Dorp, 2006, Ent. Ber. 66 (4): 125 (note,map); [AfrL]; [iNaturalist]

L. p. phlaeoides (Staudinger, 1901) Madeira

Chrysophanus phlaeas var. phlaeoides Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 74 (Knaggs in l.?)

Lycaena phlaeas phlaeoides; [CoL] ; [iNaturalist]

L. p. arethusa (Dod, 1907) Alberta

Chrysophanus arethusa Dod, 1907; Can. Ent. 39 (5): 169

Lycaena (Lycaena) hypophlaeas arethusa; [P&N23]
Lycaena (Lycaena) phlaeas arethusa; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena arethusa; [CoL]
Lycaena hypophlaeas arethusa; [iNaturalist]
Lycaena phlaeas arethusa; [NACL], #4251c

L. p. daimio (Seitz, [1909]) S.Kuriles, Sakhalin

Chrysophanus phlaeas ab. daimio Seitz, [1909]; Gross-Schmett. Erde 1: 286, pl. 77 b
Chrysophanus phlaeas daimio ab. distalba Kuwayama, 1926; Entomologist 59: 181
Lycaena phlaeus[sic] chinensis ab. kurilensis Matsumura, 1928; Insecta Matsumurana 2 (4): 200; TL: Kuriles
Chrysophanus phlaeas daimio ab. uyei Esaki & Nakahara, 1930; Zephyrus 2 (3): 150
kuriliphlaeas (Bryk, 1942) (Heodes); Dt. Ent. Z. Iris 56: 19; TL: Kurile Is.

Heodes daimio; Esaki & Nakahara, 1930, Zephyrus 2 (3): 150
Lycaena phlaeas daimio; Kudrna, 1974, Atalanta 5: 112; [BARU](note) , ; [BRU2], 124; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]; [CoL] ; [iNaturalist]
Lycaena phlaeas kuriliphlaeas; [CoL]

L. p. polaris Courvoisier, 1911 N.Urals, N.Siberia, Chukotka

Lycaena phlaeas f. polaris Courvoisier, 1911; Ent. Zs. 24 (49): 262; TL: Lappland; Norway

Chrysophanus phlaeas f. polaris; Courvoisier, 1911, Dt. ent. Z. Iris 25 (9): 109
Lycaena phlaeas polaris; [BRU2], 123; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]; [iNaturalist]

L. p. cyrenaica (Turati, 1921) Cyrenaica

Rumicia phlaeas f. cyrenaica Turati, 1921; Atti Soc. ital. Sci. nat. 60: 219
aestiva (Turati & Krüger, 1936) (Rumicia); Mem. Soc. ent. ital. 15: 57

Lycaena phlaeas cyrenaica; [CoL] ; [iNaturalist]

L. p. ethiopica (Poulton, 1922) E.Zaire (N.Kivu), SW.Uganda (Kigezi), Rwanda, Burundi, NW.Tanzania

Heodes phlaeas ethiopica Poulton, 1922; Proc. ent. Soc. Lond. 1921: lxxxvi
- = Lycaena phlaeas pseudophlaeas; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 264

Lycaena phlaeas ethiopica; [BAFR], 525; de Jong & van Dorp, 2006, Ent. Ber. 66 (4): 125 (note,map); [AfrL]; [CoL] ; [iNaturalist]

L. p. flavens (Ford, 1924) Tibet

phlaeas flavens (Ford, 1924) (Heodes); Trans. ent. Soc.: 642 [?]

Lycaena phlaeas flavens; [MRS], 663; Yoshino, 2019, Butterflies 80: 18 (note); [CoL] ; [iNaturalist]

L. p. coccineus (Ford, 1924) Tian-Shan

phlaeas coccineus (Ford, 1924) (Heodes); Trans. ent. Soc.: 726

Lycaena phlaeas coccinea; [MRS], 663
Lycaena phlaeas coccineus; [CoL] ; [iNaturalist]

L. p. japonica (Ford, 1924) Japan

phlaeas japonica (Ford, 1924) (Heodes); Trans. ent. Soc.: 729

Lycaena phlaeas japonica; [CoL] ; [iNaturalist]

L. p. indicus (Evans, 1925) Simla

Heodes phlaeas indicus Evans, 1925; J. Bombay nat. Hist. Soc. 30 (3): 616

Lycaena phlaeas indicus; Yoshino, 2019, Butterflies 80: 18 (note); [CoL] ; [iNaturalist]

L. p. lusitanicus (Bryk, 1940) Portugal

phlaeas lusitanicus (Bryk, 1940) (Heodes); Arkiv Zool. 32A (22): 20

Lycaena phlaeas lusitanicus; [CoL] ; [iNaturalist]

L. p. hibernica Goodson, 1948 Ireland

phlaeas hibernica Goodson, 1948; Entomologist 81: 177

Lycaena phlaeas hibernica; [CoL] ; [iNaturalist]

L. p. shima Gabriel, 1954 S.Arabia, Yemen

Lycaena shima Gabriel, 1954; Exped. S.W. Arabia 1937-38, 1: 388; TL: Yemen

Lycaena phlaeas shima; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 264; [BAFR], 525; [BK], 213 (note); de Jong & van Dorp, 2006, Ent. Ber. 66 (4): 125 (note,map); [AfrL]; [CoL] ; [iNaturalist]

L. p. arctodon Ferris, 1974 Montana

Lycaena phlaeas arctodon Ferris, 1974; Bull. Allyn Mus. 18: 4, f. 17-18; TL: E side Beartooth Pass, Custer NF, Carbon Co., Mont.

Lycaena (Lycaena) hypophlaeas arctodon; [P&N23]
Lycaena (Lycaena) phlaeas arctodon; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena hypophlaeas arctodon; [iNaturalist]
Lycaena phlaeas arctodon; [NACL], #4251d

L. p. kamtschatica Gorbunov, 1994? Kamchatka Peninsula

Lycaena phlaeas kamtschatica Gorbunov, 1994?; ; TL: Kamchatka, Ganal'skaya Tundra

Lycaena phlaeas kamtschatica; [iNaturalist]

L. p. ganalica Gorbunov, 1995 Kamchatka

Lycaena phlaeas ganalica; [BRU2], 123; [iNaturalist]

L. p. weberi Kohler, 2007

Lycaena (Lycaena) hypophlaeas weberi; [P&N23]
Lycaena hypophlaeas weberi; [iNaturalist]

L. p. shields Kemal & Koçak, 2007 California

Lycaena phlaeas shields Kemal & Koçak, 2007; (repl. Lycaena phlaeas alpestris Emmel & Pratt, 1998); TL: California, Mono Co., N slope of Mt. Dana
Lycaena phlaeas alpestris Emmel & Pratt, 1998; Syst. W. N. Am. Butts. (52): 662 (preocc.); TL: California, Mono Co., N slope of Mt. Dana

Lycaena (Lycaena) hypophlaeas shields; [P&N23]
Lycaena (Lycaena) phlaeas alpestris; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena hypophlaeas shields; [iNaturalist]

L. p. gonpaensis Yoshino, 2019 N.Yunnan

Lycaena phlaeas gonpaensis Yoshino, 2019; Butterflies 80: 18; TL: Shangri-La, 3300m, N.Yunnan

Lycaena kiyokoae Sakai, 1978

Afghanistan. See [maps]

Lycaena kiyokoae Sakai, 1978; Atalanta 9 (1): 115; TL: Mt Shahfuladi, 4000-4500m, Koh-i-Baba Mts, Afghanistan

Lycaena kiyokoae; [CoL]

Lycaena sichuanica Bozano & Weidenhoffer, 2001

Sichuan. See [maps]

Lycaena sichuanica Bozano & Weidenhoffer, 2001; ; TL: near Barkham, Sichuan, China

Lycaena sichuanica; Yoshino, 2019, Butterflies 80: 18 (note)

Lycaena cupreus (Edwards, 1870)

-. See [maps] Lustrous Copper

Chrysophanus cupreus Edwards, 1870; Trans. amer. ent. Soc. 3 (1): 20; TL: "Oregon"

Chalceria cupreus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 41
Lycaena (Alciphronia) cupreus cupreus; [P&N23]
Lycaena (Lycaena) cupreus; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena cuprea; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [NACL], #4252
Lycaena cupreus; [EBW]; [BOW]: pl. 20, f. 1 (text only); [NAB]; [Opler]; [MPG] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]

L. c. cupreus Oregon, ..., Wyoming

Lycaena cupreus artemisia Scott, 1981; Papilio (n.s.) 1: 6; TL: The Potholes, Teton Co., Wyoming
- = ; Pelham, 2008, J. Res. Lepid. 40: 190

Lycaena (Lycaena) cupreus cupreus; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena cupreus cupreus; [iNaturalist] ; [BOA]

L. c. snowi (Edwards, 1881) Colorado, British Columbia, Alberta, ...

Chrysophanus snowi Edwards, 1881; Trans. Kans. Acad. Sci. 7: 69; TL: Colorado
Chrysophanus snowi henryae Cadbury, 1937; Proc. Acad. Nat. Sci. Philad. 89: 409, pl. 16, f. 9; TL: Caribous Pass, B.C
- = ; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena (Heodes) snowi tr.f. mcdunnoughi Gunder, 1927; Can. Ent. 59 (12): 284, pl. A, f. 12; TL: Laggan, Alta.

Chalceria snowi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 41
Lycaena (Alciphronia) cupreus henryae; [P&N23]
Lycaena (Alciphronia) cupreus snowi; [P&N23]
Lycaena (Lycaena) cupreus snowi; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena cuprea henryae; [NACL], #4252b
Lycaena cuprea snowi; [NACL], #4252a
Lycaena cupreus henryae; ; [iNaturalist]
Lycaena cupreus snowi; [BOW]: pl. 20, f. 2; [iNaturalist] ; [BOA]
Lycaena snowi henryae; [CoL]
Lycaena snowi mcdunnoughi; [CoL]
Lycaena snowi; [CoL]
macdunnoughi[sic, mcdunnoughi]; dos Passos, 1964, Lepid. Soc. Mem. 1: 61 (emend.)

L. c. lapidicola Emmel & Pratt, 1998 California

Lycaena cuprea lapidicola Emmel & Pratt, 1998; Syst. W. N. Am. Butts. (52): 663, f. 5-8; TL: California, Mono Co., Tioga Pass
Heodes cupreus ab. ♀ maculinita Gunder, 1926; Ent. News 37 (1): 8, pl. 1, f. 11; TL: Mammoth, Mono Co., California

Lycaena (Alciphronia) cupreus lapidicola; [P&N23]
Lycaena (Lycaena) cupreus lapidicola; Pelham, 2008, J. Res. Lepid. 40: 190
Lycaena cupreus lapidicola; [iNaturalist] ; [BOA]

Lycaena helle (Denis & Schiffermüller, 1775)

CEU, NEU, Russia, Siberia, Amurland, Dzhungarsky Alatau. See [maps] Czerwończyk fioletek Blauschillernder Feuerfalter Luhtakultasiipi Anilinvinge Violet Copper

Papilio helle Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 181
Papilio amphidamas Esper, 1781; Die Schmett. Th. I, Bd. 2 (3): 82, (2): pl. 58, f. 4, (3): pl. 63, f. 5
lapponica (Baukhaus, 1876) (Chrysophanus); Entom. Mon. 1: 40; TL: Lappland
Polyommatus amphidammas ab. obscura Lampa, 1885; Ent. Tidskr. 6 (1): 13
obscura (Rühl, 1892) (Polyommatus); Palaearkt. Grossschm. 1: 220
aestiva (Wahlgren, 1924) (Heodes); Ent. Tidskr. 45: 97 (?repl. Polyommatus amphidamas ab. obscura Lampa, 1885)
minor (Derenne, 1925) (Chrysophanus); Rev. Mens. Soc. ent. Namur. 25: 92
marchica (Hannemann, 1928) (Chrysophanus); Int. ent. Zs. 22: 210
Heodes amphidamas pyrenaica Deslandes, 1930; Bull. Soc. ent. Fr. 35 (15): 243; TL: Porté (Pyr.-Orientales), 1600m
ferephlaeas (Hannemann, 1931) (Chrysophanus); Int. Ent. Zs. 24 (37/38): 401; TL: Germany
fulva (Hannemann, 1931) (Chrysophanus); Int. Ent. Zs. 24 (37/38): 401; TL: Germany
radiata (Hannemann, 1931) (Chrysophanus); Int. Ent. Zs. 24 (37/38): 401; TL: Germany
deslandesi Hemming, 1932; Proc. ent. Soc. Lond. 7: 29 (repl. pyrenaica Deslandes, 1930); TL: Pyrenees
leonia Beuret, 1936; Lambillionea 36: 272; TL: Bernese Jura
striata (Hackray, 1938) (Heodes); Lambillionea 38: 20
yamashinai (Matsumura, 1939) (Chrysophanus); Bull. biogeogr. Soc. Japan 9 (20): 356
radiata (Valle, 1945) (Chrysophanus); Ann. ent. Fenn. 7: 205; TL: Finland
arvernica Bernardi & de Lesse, 1952; Rev. franc. Lép. 13: 209
hellesimilis (Beuret, 1952) (Helleia); Mitt. ent. Ges. Basel (n.f.) 2: 101; TL: Switzerland
antelutea Beuret, 1954; Lycaeniden Schweiz 1: 57; TL: Switzerland
bicolor Beuret, 1954; Lycaeniden Schweiz 1: 57; TL: Switzerland
caeruleolunulata Beuret, 1954; Lycaeniden Schweiz 1: 57; TL: Switzerland
juragloria Beuret, 1954; Lycaeniden Schweiz 1: 58; TL: Switzerland
reducta Beuret, 1954; Lycaeniden Schweiz 1: 58; TL: Switzerland
tristis Beuret, 1954; Lycaeniden Schweiz 1: 58; TL: Switzerland
magdalenae Guérin, 1959; Alexanor 1: 88; TL: France
eneli Betti, 1977; Alexanor 10 (2): 92; TL: France
perrettei Weiss, 1977; Linneana Belg. 6 (10): 255

Helleia helle antelutea; [CoL]
Helleia helle arvernica; [CoL]
Helleia helle bicolor; [CoL]
Helleia helle caeruleolunulata; [CoL]
Helleia helle deslandesi; [CoL]
Helleia helle eneli; [CoL]
Helleia helle hellesimilis; [CoL]
Helleia helle juragloria; [CoL]
Helleia helle lapponica; [CoL]
Helleia helle leonia; [CoL]
Helleia helle magdalenae; [CoL]
Helleia helle marchica; [CoL]
Helleia helle perrettei; [CoL]
Helleia helle reducta; [CoL]
Helleia helle tristis; [CoL]
Helleia helle yamashinai; [CoL]
Helleia helle; [MRS], 662; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [CoL]
Lycaena (Lycaena) helle; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; [RFEL]
Lycaena helle magdalenae; [iNaturalist]
Lycaena helle; [SPRK]; [BARU], #342; [H&R]; [EBW]; [BOW]: pl. 9, f. 37; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BRU2]: 123, pl. 57, f. 1-6; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF] ; [Mario Maier]

650x654 (~39Kb) ♀ FINLAND: Ks: Kuusamo Liikasenvaara, 736:61, 9.7.1998, Photo © Markku Savela

500x626 (~29Kb) ♂ FINLAND: Ks: Kuusamo Liikasenvaara, 736:61, 9.7.1998, Photo © Markku Savela

1000x1000 (~197Kb) ♂ upperside Russia, West Siberia, Novosibirsk Province, Toguchin District, the Salairskii Kryazh Range, the Suenga River valley, an opening in fir taiga, 6th June 2000 (on Euphorbia lutescens), Photo © Oleg Kosterin

L. h. helle CEU, NEU, Caucasus Major

Lycaena helle helle; [BRU2], 123; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74

L. h. phintonis (Fruhstorfer, 1910) Amurland

Chrysophanus amphidamas phintonis Fruhstorfer, 1910; Ent. Zs. 24 (26): 144; TL: Sibirien, Irkutsk

Helleia helle phintonis; [CoL]
Lycaena helle phintonis; [BARU](note) , ; [BRU2], 123; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74; [RFEL]

600x835 (~115Kb) upperside underside Russia: S.Tuva, Korumnug-Taiga Mts. (1200m), 8-25.6.2002, Vashchenko A. & B. leg. Photo © D. Smirnov

L. h. shchurovi Gorbunov, 2001 Caucasus

Lycaena helle shchurovi Gorbunov, 2001; Butt. Russ. classif.: 110; TL: Lagonaki Plateau, Caucasus

Lycaena helle shchurovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 74

1287x903 (~184Kb) underside Russian Federation, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, tall herbage at fir forest margin, 13 July 2008, Photo © Oleg Kosterin

Subgenus Thersamonia Verity, 1919

Thersamonia Verity, 1919; Ent. Rec. J. Var. 31: 28; TS: Papilio thersamon Esper
- = Lycaena; [NL4A], 138
- = Subgenus Heodes; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)

Subgenus Thersamonia (Lycaena); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia; [CoL] ; [NHM-BMW]
Thersamonia (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
Thersamonia (Lycaenidae); [BRU2], 127

Lycaena thersamon (Esper, 1784)

it, SEEU, Asia Minor, W.Asia, Iraq, Iran, N.Altai, ?Mongolia, NW.China. See [maps] Lesser Fiery Copper

Papilio thersamon Esper, 1784; Die Schmett. Th. I, Bd. 2 (9): 176, pl. 89, f. 6; TL: Sarepta, Südrussland
omphale Klug, 1834; in Ehrenberg, Symbolae phys., Ins. 4: pl. 40, f. 12-14
?hylla Koçak, 1983; (preocc.)

Lycaena (Thersamonia) thersamon; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena thersamon; [MRS], 663; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE]
Polyommatus thersamon; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 363
Thersamonia thersamon; Verity, 1919, Ent. Rec. J. Var. 31: 28; [H&R]; [BARU], #344; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 10; [EBW]; [BMAT]: 106 (Oberthür 1915); [BOW]: pl. 11, f. 3; [BRU2]: 127, pl. 59, f. 1-12; [CoL] ; [Mario Maier]

500x398 (~97Kb) upperside Hisaronu limani, SW Turkey 18/04/95, Photo © Simon Coombes

500x444 (~68Kb) underside Hisaronu limani, SW Turkey 18/04/95, Photo © Simon Coombes

L. t. thersamon

Lycaena thersamon thersamon; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia thersamon thersamon; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 10

L. t. persica (Bienert, [1870]) Kopet-Dagh, Ghissar

Polyommatus thersamon var. persica Bienert, [1870]; Lepid. Erg. Reise Persiens: 28; TL: Persia

Lycaena thersamon persica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia thersamon persica; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 12; [BRU2], 127; [CoL]

L. t. jiadengyuensis (Huang & Murayama, 1992) Xinjiang (Altai)

Thersamonia thersamon jiadengyuensis Huang & Murayama, 1992; Tyô to Ga 43 (1): 9, f. 27-28; TL: Jiadengyu, Altai, 1400m
- = Lycaena thersamon sajanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75

L. t. sajanica (Huang & Murayama, 1992) Sajan Mts.

Thersamonia thersamon sajanica Huang & Murayama, 1992; Tyô to Ga 43 (1): 11; TL: Turan, Sajan Mts., 2000m, SW.Irkutsk

Lycaena thersamon sajanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75

Lycaena kurdistanica (Riley, 1921)

Armenia (highland), NE.Turkey, W.Iran. See [maps]

Heodes thersamon kurdistanica Riley, 1921; Ann. Mag. nat. Hist. (9) 8 (47): 598; TL: Harir, Karind Gorge; Kermanshah

Lycaena (Thersamonia) kurdistanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena kurdistanica; [iNaturalist] ; [LEPIFORUM]
Thersamonia kurdistanica; [BRU2]: 128, pl. 59, f. 13-15
Thersamonia thersamon kurdistanica; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 11; [CoL]

Lycaena alaica (Grum-Grshimailo, 1888)

Pamirs-Alai, S.Ghissar, NE.Afghanistan?. See [maps]

Polyommatus thersamon var. alaica Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4): 305; TL: Alai [Malyi Karamyk River, sothern slope o fthe Alai Mts. near Daraut-Kurgan, Kirghizia]

Lycaena (Thersamonia) alaica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena alaica;
Lycaena thersamon alaica; [MRS], 663
Polyommatus thersamon var. alaica; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 364
Thersamonia alaica; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 14; [BRU2]: 129, pl. 59, f. 16-18

Lycaena phoebus (Blachier, 1905)

Morocco. See [maps] Moroccan Copper

Chrysophanus phoebus Blachier, 1905; Bull. Soc. ent. Fr. 1905: 212; TL: Atlas Mountains (Morocco)
Chrysophanus phoebus ab. aurata Lucas, 1937; Bull. Soc. ent. Fr. 42 (16): 233; TL: Marrakech (Morocco)

Lycaena (Thersamonia) phoebus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name)
Lycaena phoebus; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE]
Thersamonia phoebus; [H&R]; [EBW]; [BOW]: pl. 11, f. 3 (text only); Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 15; [BMAT], 29, pl. 10, f. 16-21

600x435 (~36Kb) upperside Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Michel Tarrier	600x435 (~33Kb) underside Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Jean Delacre	600x435 (~38Kb) Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Jean Delacre
600x435 (~44Kb) underside Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Jean Delacre

Lycaena lampon (Lederer, [1870])

Kopet-Dagh, SE.Turkey, Iran, Afghanistan. See [maps]

Polyommatus lampon Lederer, [1870]; Horae Soc. ent. ross. 8 (1): 8, pl. 2; TL: Gebirge Lendakuh; Hadschyabad

Heodes lampon; [BOW]: pl. 205, f. 35 (text)
Lycaena (Thersamonia) lampon; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena lampon; [CoL] ; [iNaturalist] ; [LEPIFORUM]
Thersamonia lampon; Verity, 1919, Ent. Rec. J. Var. 31: 28; [BOW]: pl. 11, f. 4 (text); Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 28; [BRU2]: 129, pl. 59, f. 28-30; [CoL]

L. l. lampon

Thersamonia lampon lampon; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 29

L. l. eberti Forster, 1972 Afghanistan

Lycaena lampon eberti Forster, 1972; Ent. Zs. 82 (20): 225; TL: Afghanistan, Band-i-amir

Lycaena lampon eberti; [CoL]
Thersamonia eberti; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 35

Lycaena lamponides (Staudinger, 1901)

Chrysophanus lampon v. lamponides Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 73

Lycaena (Thersamonia) lamponides; ten Hagen & Eckweiler, 2000, Nachr. Ent. Ver. Apollo NF 21 (2): (49-54)
Lycaena lamponides; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Lycaena aditya (Moore, [1875])

W.Pamirs, Afghanistan, Pakistan, N.India. See [maps]

Chrysophanus aditya Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 571, pl. 66, f. 1; TL: Dros valey, Ladak [N.Pakistan]

Lycaena (Thersamonia) aditya; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena aditya; [CoL] ; [iNaturalist]
Thersamonia aditya; [BRU2]: 130, pl. 59, f. 40-42
Thersamonia solskyi aditya; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 39

L. a. charmesse Lamas, 2008

Lycaena aditya charmesse Lamas, 2008; Zootaxa 1848: 49 (repl. Lycaena aditya churkini Charmeux & Desse, 2006)
Lycaena aditya churkini Charmeux & Desse, 2006; 85, 89, pl. 3, f. 1-3, 7-9 (preocc. Athamanthia churkini Zhdanko, 2000)
- = ; Lamas, 2008, Zootaxa 1848: 49

Lycaena solskyi Erschoff, 1874

Pamirs-Alai, Ghissar-Darvaz, Tian-Shan, Kashmir, Samarkand. See [maps]

Lycaena solskyi Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5): 8, pl. 1, f. 7; TL: Maracanda [Uzbekistan]

Heodes solskyi; [BOW]: pl. 205, f. 34, pl. 206, f. 1
Lycaena (Thersamonia) solskyi; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena solskyi; [MRS], 664; [iNaturalist] ; [LEPIFORUM]
Polyommatus solskyi thersamon; [CoL]
Polyommatus solskyi; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 358; [CoL]
Thersamonia solskyi; [EBW], (name); Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 38; [BRU2]: 129, pl. 59, f. 31-36

L. s. solskyi Ghissarsky, Turkestansky, Zeravshansky Mts.

Lycaena solskyi solskyi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]
Thersamonia solskyi solskyi; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 39; [BRU2], 129

L. s. fulminans (Grum-Grshimailo, 1888) Zaalaisky Mts, Tian-Shan, Alai, Darvaz

Polyommatus solskyi var. fulminans Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4): 305; TL: Translai

Lycaena solskyi fulminans; [MRS], 664; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]
Polyommatus solskyi fulminans; [CoL]
Polyommatus solskyi var. fulminans; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 358, pl. 6, f. 4a-b
Thersamonia solskyi fulminans; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 40; [BRU2], 129

L. s. fulmineus (Turati, 1933) Himalayas

solskyi fulmineus (Turati, 1933) (Chrysophanus); Boll. Soc. ent. Ital. Sci. nat. 65 (1): 13

Polyommatus solskyi fulmineus; [CoL]

L. s. attila (Zhdanko, 1990) Zaalaisky Mts.

Thersamonia attila Zhdanko, 1990; Ent. Obozr. 69 (1): 140; TL: [Irkeshtam, 2600m, eastern part of Zaalaisky Mts., Kirghizia]

Lycaena attila;
Lycaena solskyi attila; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia attila; [BRU2]: 130, pl. 59, f. 43, pl. 83, f. 43

Lycaena alpherakii (Grum-Grshimailo, 1888)

E.Pamirs, NW.China. See [maps]

Polyommatus alpherakii Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4): 305; TL: Hindukush orient.

Lycaen (Thersamonia) alpherakii; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena alpherakyi[sic];
Polyommatus alpherakii; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 359, pl. 14, f. 2
Polyommatus solskyi alpherakii; [CoL]
Thersamonia alpherakyi[sic, alpherakii]; [BRU2]: 130, pl. 59, f. 37-39
Thersamonia solskyi alpherakii; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 40
Thersamonia solskyi alpherakyi[sic];

L. a. alpherakii

Lycaena alpherakii alpherakii; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75

L. a. zhdankoi (Churkin, 2002) W.Pamirs

Thersamonia alpherakii zhdankoi Churkin, 2002; Helios 3: 183, pl. 15, f. 16-20 (?preocc. Athamanthia zhdankoi Lukhtanov, 2000); TL: Tadzhikistan, Pamirs, Vanchsky mst, Gushkon valley, 3000-3300m
- = Lycaena alpherakii sergeii; Lamas, 2008, Zootaxa 1848: 49
Lycaena alpherakii sergeii Lamas, 2008; Zootaxa 1848: 49 (repl. Thersamonia alpherakii zhdankoi Churkin, 2002)

Lycaena alpherakii zhdankoi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75

Lycaena asabinus (Gerhard, 1850)

Lebanon - Turkey - Transcaucasia, Iran, Armenia. See [maps]

Polyommatus asabinus Gerhard, 1850; Versuch Mon. europ. Schmett. (3): 7, pl. 9, f. 3a-c
Polyommatus asabinus Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 1 (49): (ii) pl. 109, f. 527-528; TL: Amasya, Turkey
Polyommatus helius Herrich-Schäffer, [1852]; Syst. Bearb. Schmett. Europ. 6 (55): 32
- = Thersamonia asabinus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 43

Lycaena (Thersamonia) asabinus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena asabinus; [CoL] ; [iNaturalist] ; [LEPIFORUM]
Thersamonia asabinus; [EBW]; [BOW]: pl. 11, f. 3 (text only); Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 43; [BRU2]: 128, pl. 59, f. 22-24; [CoL]
Thersamonia asebinus[sic, asabinus]; Verity, 1919, Ent. Rec. J. Var. 31: 28

1124x1162 (~321Kb) ♂ underside Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

660x726 (~143Kb) ♂ underside Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

1079x1088 (~329Kb) ♂ underside Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

L. a. asabinus

Thersamonia asabinus asabinus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 43

L. a. satraps (Staudinger, 1878)

Polyommatus satraps Staudinger, 1878; Horae Soc. Ent. Ross. 14: 230; TL: Külek (Taurus)

Thersamonia asabinus satraps; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 44
Thersamonia satraps; Verity, 1919, Ent. Rec. J. Var. 31: 28

L. a. tauricus (Heyne, [1895])

var. tauricus Heyne, [1895]²; Palaearkt. Grossschm. 1: 744; TL: Taurus

Thersamonia asabinus tauricus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 44

Lycaena ochimus (Herrich-Schäffer, [1851])

Transcaucasia, Turkey, Syria, Lebanon. See [maps]

Polyommatus ochimus Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 1 (49): (ii) f. 523-526; TL: Kleinasien
Polyommatus ochimus Herrich-Schäffer, [1852]; Syst. Bearb. Schmett. Europ. 6 (55): 31
phaeton (Freyer, [1850]) (Polyommatus); Neuere Beitr. Schmett. 6: pl. 571, f. 2 (preocc.)
Chrysophanus ochimus ab. strigifera Schultz, 1905; Ent. Zs. 18 (33): 131
Polyommatus kefersteini Gerhard, 1850; Versuch Mon. europ. Schmett. (3): 7, pl. 9, f. 4a-c (nom. nud.)
- = Thersamonia ochimus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 20

Heodes ochimus; [BOW]: pl. 205, f. 34
Lycaena (Thersamonia) ochimus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena kefersteinii; [iNaturalist]
Lycaena ochimus; [iNaturalist] ; [LEPIFORUM]
Thersamonia ochimus; [EBW]; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 20; [BRU2]: 128, pl. 59, f. 19-21; [CoL]

L. o. ochimus Armenia (highland)

Lycaena ochimus ochimus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia ochimus ochimus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 20; [BRU2], 128

L. o. ocliferius (Nekrutenko, 1977) Caucasus Minor

Heodes (Thersamonia) ochimus ocliferius Nekrutenko, 1977; Rep. Acad. Sci. SSR Ukraine (B) 2: 457

Lycaena ochimus ocliferius; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia ochimus ocliferius; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 21; [BRU2], 128; [CoL]

L. o. sahandensis (Schurian & Hofmann, 1982) Talysh Mts.

Thersamonia ochimus sahandensis Schurian & Hofmann, 1982; Nachr. Ent. Ver. Apollo Suppl. 2: 22; TL: 25 m. S. E. of Tabriz, Iran, Gombar, Sahand Dagh Mtns, 8500-10500ft

Lycaena ochimus sahandensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Thersamonia ochimus sahandensis; [BRU2], 128

Lycaena thetis Klug, 1834

Greece, Asia Minor - Armenia (highlands), Iraq, Iran, Baluchistan, Chitral, Ladak. See [maps] Golden Copper Fiery Copper

Lycaena thetis Klug, 1834; in Ehrenberg, Symbolae phys., Ins. 4: pl. 40, f. 17-18; TL: Syria
Chrysophanus caudatus Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 73
Chrysophanus thetis var. zahaltensis Graves, 1910; Ent. Rec. J. Var. 22: 273; TL: Syria, "Ain Zahalta Cedars"

Lycaena (Thersamonia) thetis; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena thetis caudatus; [CoL] ; [iNaturalist]
Lycaena thetis thetis; [iNaturalist]
Lycaena thetis; [BIR], 302; [Otakar Kudrna]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE]
Thersamonia thetis; [EBW]; [H&R]; [BOW]: pl. 11, f. 4; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 48; [BRU2]: 129, pl. 59, f. 25-27

L. t. ignitus (Herrich-Schäffer, [1846])

Polyommatus ignitus Herrich-Schäffer, [1846]; Syst. Bearb. Schmett. Europ. 1 (14): (ii) f. 332

Thersamonia thetis ignitus; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 50

L. t. elbursina (Pfeiffer, 1937)

Chrysophanus thetis elbursina Pfeiffer, 1937; Mitt. Münch. Ent. Ges. 27 (1): 36; TL: Elburs, Tacht i Suleiman

Lycaena thetis elbursina; [CoL]
Thersamonia thetis elbursina; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 50

Lycaena hyrcana (Neuburger, 1903)

N.Iran - C.Asia. See [maps]

Chrysophanus ochimus var. hyrcana Neuburger, 1903; Soc. Ent. 18 (8): 59; TL: Hyrcania

Thersamonia ochimus hyrcana; Schurian & Hofmann, 1982, Nachr. Ent. Ver. Apollo Suppl. 2: 25; [CoL]

Subgenus Thersamolycaena Verity, 1957

Thersamolycaena Verity, 1957; Ent. Rec. J. Var. 69: 225 (repl. Disparia Verity, 1943); TS: Papilio dispar Haworth
Subgenus Disparia Verity, 1943 (Lycaena); Le Farfalle diurn. d'Italia 2: 21, 58 (preocc. Disparia Nagano, 1916); TS: Papilio dispar Haworth
- = Lycaena; [NL4A], 138
Subgenus Rapsidia Sibatani, 1974 (Lycaena); Aust. ent. Soc. 1974 (13): 109 (repl. Disparia Verity, 1943); TS: Papilio dispar Haworth
Subgenus Alciphronia Koçak, 1992 (Heodes); ; TS: Papilio alciphron Rottemburg
- = Subgenus Heodes; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Subgenus Mirzakhania Koçak, 1996 (Thersamolycaena); ; TS: Chrysophanus kasyapa Moore
- = Subgenus Heodes; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)

Subgenus Thersamolycaena (Lycaena); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Alciphronia; [iNaturalist] ; [NHM-BMW]
Lycaena (Alciphronia); [P&N23]
Mirzakhania; [NHM-BMW]
Rapsidia; [NHM-BMW]
Thersamolycaena; [NHM-BMW]
Thersamolycaena (Lycaenidae); [BRU2], 125

Lycaena dispar (Haworth, 1802)

. See [maps] Czerwończyk nieparek Isokultasiipi Dukatvinge Large Copper Großer Feuerfalter

Papilio dispar Haworth, 1802; Prodr. Lep. Brit.: 3, nota; TL: England
Papilio hippothoe Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 181
Papilio hippothoe Esper, 1778; Die Schmett. Th. I, Bd. 1 (9): 350, (7): pl. 38, f. 1a-b
Chrysophanus dispar ab. nigrolineata Verity, 1904; Entomologist 37 (490): 57; TL: Modena
Chrysophanus dispar ab. cuneigera Tutt, 1906; Nat. Hist. Br. Lepid. 8: 428
Chrysophanus dispar ab. subcuprea Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. transversa Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. parva Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. excessa Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. subobsoleta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. obsoleta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 430
Chrysophanus dispar ab. nigrescens Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. neurata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. cuneata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. bilineata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. unilineata Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. suppressa Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. minor Tutt, 1906; Nat. Hist. Br. Lepid. 8: 431
Chrysophanus dispar ab. brunnescens Tutt, 1906; Nat. Hist. Br. Lepid. 8: 432
Chrysophanus dispar ab. punctijuncta Tutt, 1906; Nat. Hist. Br. Lepid. 8: 432
Chrysophanus dispar ab. pupurpurascens Tutt, 1906; Nat. Hist. Br. Lepid. 8: 432
Chrysophanus rutilus ab. radiata Courvoisier, 1907; Zs. wiss. Insektenbiol. 3 (2): 37
Chrysophanus dispar var. rutilus ab. aurata Hirschke, 1910; Verh. zool.-bot. Wien 60: 412
rutilus ab. basi-novopuncta Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 58
rutilus ab. paucipuncta Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 63
Chrysophanus dispar var. rutilus ab. heyni Strand, 1913; Ent. Rdsch. 30: 98
Chrysophanus dispar var. burdigalensis Lucas, 1913; Int. Ent. Zs. 6: 282; TL: Bordeaux
Chrysophanus dispar rutilus f. extincta Hensel, 1920; Int. ent. Zs. 14 (16): 122; TL: Berlin
obscurior (Pionneau, 1937) (Chrysophanus); Echange 53: 3; TL: France
carueli Le Moult, 1945; Misc. Ent. 42: 55
postcarueli Le Moult, 1945; Misc. Ent. 42: 56
verityi Le Moult, 1945; Misc. Ent. 42: 57
Thersamonia dispar r. centralitaliae Verity, 1946; Le Farfalle diurn. d'Italia 2: 61
Thersamonia dispar r. centralitaliae f. italiae Verity, 1946; Le Farfalle diurn. d'Italia 2: 61
Thersamonia dispar f. ignita Lempke, 1954; Tijdschr. Ent. 97: 336
Thersamonia dispar f. latefasciata Lempke, 1954; Tijdschr. Ent. 97: 336
Thersamonia dispar f. antinigrescens Lempke, 1954; Tijdschr. Ent. 97: 336
Thersamonia dispar f. ♂ postnigricosta Lempke, 1954; Tijdschr. Ent. 97: 336
Thersamonia dispar f. supradiscoelongata Lempke, 1954; Tijdschr. Ent. 97: 337
Thersamonia dispar f. ♂ ornata Lempke, 1954; Tijdschr. Ent. 97: 337
Thersamonia dispar f. ♂ postdisconulla Lempke, 1954; Tijdschr. Ent. 97: 337
Thersamonia dispar f. centroelongata Lempke, 1954; Tijdschr. Ent. 97: 337
Thersamonia dispar f. infraparvipuncta Lempke, 1954; Tijdschr. Ent. 97: 338
depuncta (Beuret, 1954) (Thersamonia); Lycaeniden Schweiz 1: 77
decolorata (Beuret, 1954) (Thersamonia); Lycaeniden Schweiz 1: 79
gronieri Bernardi, 1963; Alexanor 3: 53; TL: France

Lycaena (Rapsidia) dispar; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena (Thersamolycaena) dispar; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]
Lycaena dispar antinigrescens; [CoL]
Lycaena dispar burdigalensis; [CoL]
Lycaena dispar carueli; [CoL] ; [iNaturalist]
Lycaena dispar centralitaliae; [CoL]
Lycaena dispar centroelongata; [CoL]
Lycaena dispar decolorata; [CoL]
Lycaena dispar depuncta; [CoL]
Lycaena dispar extincta; [CoL]
Lycaena dispar gronieri; [CoL]
Lycaena dispar hippothoe; [CoL] ; [iNaturalist]
Lycaena dispar ignita; [CoL]
Lycaena dispar infraparvipuncta; [CoL]
Lycaena dispar italiae; [CoL]
Lycaena dispar latefasciata; [CoL]
Lycaena dispar ornata; [CoL]
Lycaena dispar postdisconulla; [CoL]
Lycaena dispar postnigricosta; [CoL]
Lycaena dispar supradiscoelongata; [CoL]
Lycaena dispar verityi; [CoL]
Lycaena dispar; [H&R]; [EBW]; [BOW]: pl. 9, f. 36, 38; [MRS], 663; [Otakar Kudrna]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF] ; [Mario Maier]
Thersamolycaena dispar; [BARU], #346; [BRU2]: 125, pl. 58, f. 7-12
Thersamonia dispar;

597x400 (~64Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	594x400 (~45Kb) ♂ upperside Central Europe 1990-2000, Photo © Michael Borsch	470x364 (~80Kb) ♂ upperside Czech Republic, Blansko, 4.7.2004, Photo © Michal Koupý
470x434 (~85Kb) underside Czech Republic, Blansko, 4.VII.2004, Photo © Michal Koupý	470x319 (~89Kb) ♀ upperside Czech Republic, Blansko, 17.7.2004, Photo © Michal Koupý

L. d. dispar England (extinct)

Thersamolycaena dispar dispar; [BRU2], 126

L. d. rutilus (Werneburg, 1864) Europe, Caucasus, Transcaucasia, N.Tien-Shan, W.Tien-Shan, Dzhungarsky Alatau, Ghissar

Papilio rutilus Werneburg, 1864; Beir. zur Schmet. (1): 391
Polyommatus dispar v. rutilus ab. sagittifera Hormuzaki, 1893; Soc. Ent. 8: 58
Polyommatus dispar var. (gen. I) vernalis Hormuzaki, 1893; Soc. Ent. 8: 130
Chrysophanus dispar rutilus f. continentalis Courvoisier, 1912; Int. ent. Zs. 6 (10): 65
Chrysophanus dispar var. rutilus ab. gabori Diószeghy, 1913; Rovartani Lapok 20: 193
Chrysophanus dispar var. rutilus ab. bimaculatus Diószeghy, 1913; Rovartani Lapok 20: 194
Chrysophanus dispar var. rutilus ab. minor Diószeghy, 1913; Rovartani Lapok 20: 194
Chrysophanus dispar v. rutilus f. obscurior Hoffmann, 1914; Mitt. Nat. Ver. Steierm. 50: 288
Chrysophanus dispar rutilus ab. ♀ semibrunnea Müller, 1920; Int. ent. Zs. 14 (14): 108
Chrysophanus dispar rutilus (gen. vern.) ab. albidoflava Bayer, 1924; Verh. zool.-bot. Ges. Wien 73 (S.B.): 21
fulgurata (Mezger, 1931) (Chrysophanus); Lambillionea 31: 23
brunnea (Mezger, 1934) (Heodes); Lambillionea 34: 98
punctata (Mezger, 1934) (Heodes); Lambillionea 34: 98
Lycaena dispar hungarica Szabó, 1956; Folia ent. hung. (n.s.) 9: 302, 340
- = Lycaena dispar rutila; Bálint, 1990, Janus Pannonius Múz. 34: 49

Lycaena dispar hungarica; [CoL]
Lycaena dispar rutila; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]
Lycaena dispar rutilus; [CoL] ; [iNaturalist] ; [Heiner Ziegler]
Thersamolycaena dispar rutila; [Oleg Kosterin]; [BRU2], 126; Yakovlev, 2012, Nota lepid. 35 (1): 67

533x400 (~47Kb) ♀ upperside The Black Sea coast at the settlement Yuzhnyaya Ozereevka near Novorossiysk, Novorossiysk District, Krasnodarskii Krai Province, Russia. 2nd June 1990, Photo © Oleg Kosterin

1210x1024 (~304Kb) ♂ Russian Federation, Krasnodarskii Krai, Adler, bank of Nizhneimeretinskaya harbour, 15 July 2008, Photo © Oleg Kosterin

L. d. aurata Leech, 1887 C.Siberia, E.Amur, Ussuri

Polyommatus auratus Leech, 1887; Proc. zool. Soc. Lond. 1887: 414, pl. 35, f. 3; TL: Korea
Polyommatus dispar var. borodowskyi Grum-Grshimailo, 1899; Ann. Mus. Zool. Imp. Acad. Sci. St.-Petersb. 4 (4): 459
Chrysophanus dispar parvus Kurentzov, 1941

Lycaena dispar aurata; [MRS], 663; [iNaturalist]
Lycaena dispar auratus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]
Lycaena dispar borodowskyi; [MRS], 663; [CoL]
Polyommatus auratus; [CoL]
Thersamolycaena dispar aurata; [BARU](note) , ; [BRU2], 126

533x400 (~36Kb) ♀ underside A road crossing a woody (Larix gmelinii (Rupr.) Rupr.) brook valley between the villages Budyumkan and Uryupino 5-7 km of the former, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 25th July 1997, Photo © Oleg Kosterin

L. d. dahurica (Graeser, 1888) Transbaikalia, W.Amur

Polyommatus dispar var. dahurica Graeser, 1888; Berl. Ent. Zs. 32 (1): 75

Lycaena dispar dahurica; [CoL]
Thersamolycaena dispar dahurica; [BARU](note) , ; [BRU2], 126

L. d. festiva (Krulikovsky, 1909) Ural, W.Siberia

Chrysophanus dispar festivus Krulikovsky, 1909; Rev. russ. ent. 9 (3): 300
ramaini (Le Moult, 1946) (Heodes); Novit. Ent. 11-12: 141
primus (Le Moult, 1946) (Heodes); Novit. Ent. 11-12: 141
subauratus (Le Moult, 1946) (Heodes); Novit. Ent. 11-12: 141

Lycaena dispar festivus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena dispar ramaini; [CoL]
Lycaena primus; [iNaturalist]
Thersamolycaena dispar festiva; [BARU](note) , ; [BRU2], 126

457x400 (~41Kb) ♀ underside The Irtysh River left bank floodland within the Omsk City, West Siberia, Russia. 27th August 1996 (on Inula salicina L.), Photo © Oleg Kosterin

L. d. batavus Oberthäur, 1923

Chrysophanus batavus Oberthür, 1923; Etud. Lépid. Comp. 21 (1): 73; TL: Holland
posticeoatrata (Mezger, 1931) (Chrysophanus); Lambillionea 31: 22

Lycaena dispar batava; ; [iNaturalist]
Lycaena dispar batavus; [CoL]

Lycaena alciphron (Rottemburg, 1775)

SEU, CEU, Asia Minor, Iran, W.Sibera, Altai, E.Kazakhstan, Zabaikalye, Mongolia, N.Sikhote-Alin. See [maps] Czerwończyk zamgleniec Manto de Purporo Violetter Feuerfalter Purple-shot Copper

Papilio alciphron Rottemburg, 1775; Der Naturforscher 6: 11; TL: Berlin
Papilio virgaurea Hufnagel, 1766; Berlin. Mag. 2: 79 (preocc.)
Papilio lampetie Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 322
Papilio hipponoe Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): 68, pl. 62, f. 2, (5): pl. 78, f. 5-6
Papilio hiere Fabricius, 1787; Mantissa Insectorum 2: 80
Papilio helle Borkhausen, 1788; Naturges. Eur. Schmett. 1: 146
Polyommatus alciphron intermedia Stefanelli, 1874; Bull. Soc. ent. ital. 6: 85; TL: Tuscany
Chrysophanus alciphron ab. constricta Schultz, 1900; Ill. Zs. Ent. 5: 234
Chrysophanus alciphron ab. ♀ midas Wheeler, 1903; Butts. Switzerland: 16; TL: Martigny
Chrysophanus alciphron ab. mutilata Schultz, 1905; Ent. Zs. 18 (33): 131; TL: Berlin
Chrysophanus alciphron ab. ♂ viduata Schultz, 1905; Ent. Zs. 18 (34): 133
Chrysophanus alciphron ab. ♀ infulvata Schultz, 1905; Ent. Zs. 18 (34): 133
Chrysophanus alciphron ab. caerulescens Rebel, 1910; in Berge, Schmetterlingsbuch (edn 9): 62; TL: Baden by Wien
Polyommatus gordius albescens Oberthür, 1910; Ètud. Lépid. Comp. 4: 114, 667, pl. 38, f. 247; TL: Graubunden
Polyommatus gordius nevadensis Oberthür, 1910; Ètud. Lépid. Comp. 4: 114, 667, pl. 38, f. 248; TL: Sierra Nevada
Polyommatus gordius rondoui Oberthür, 1910; Ètud. Lépid. Comp. 4: 114, 667, pl. 38, f. 246
Polyommatus gordius bellieri Oberthür, 1910; Ètud. Lépid. Comp. 4: 114, 667, pl. 38, f. 249-250; TL: Sicily
Chrysophanus alciphron ab. gerhardti Hirschke, 1910; Verh. zool.-bot. Wien 60: 412
gordius[?] confluentia multiplex [= confluentiamultiplex] Courvoisier, 1911; Ent. Zs. 24 (47): 255
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 115
Chrysophanus alciphron aetnea Turati, 1911; Soc. Ent. 25 (21): 81, 82; TL: Nicolosi, Etnea
Chrysophanus alciphron ruehli Turati, 1911; Soc. Ent. 25 (21): 83; TL: Abruzzo
Chrysophanus alciphron columbanus f. mediteranea Turati, 1911; Soc. Ent. 25 (21): 84; TL: Ceriana
violacea (Ksienschopolski, 1911) (Chrysophanua); Trudy obshch.Izsl. Volyni Zitomir 8: 55
bi-centrojuncta [= bicentrojuncta] Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 51
basi-novopuncta [= basinovopuncta] Courvoisier, 1912; Dt. ent. Z. Iris 26 (1): 58
Chrysophanus alciphron ab. ♂ pallida Fritsch, 1912; Ent. Rundsch. 29 (21): 135; TL: Liegnitz
Loweia alciphron r. mirabilis Verity, 1919; Ent. Rec. J. Var. 31: 28
Loweia alciphron insignis de Sagarra, 1924; Butll. Inst. Catal. Hist. nat. (2) 4 (9): 201; TL: Albarracin (Aragó), 1100m
Loweia alciphron paupercula de Sagarra, 1924; Butll. Inst. Catal. Hist. nat. (2) 4 (9): 201; TL: Puerto de Orihuela del Tremedal (Aragó), 1700m
Loweia alciphron r. ultragordius Verity, 1926; Ent. Rec. J. Var. 38: 105; TL: Cottian Alps
Chrysophanus alciphron f. bellieri ab. citrina Züllich, 1929; Zs. Öst. EntVer. 14 (5): 52
cecilia (Pionneau, 1936) (Chrysophanus); Echange 52: 24
emarginata (Warnecke, 1942) (Chrysophanus); Dt. ent. Z. Iris 56: 103; TL: Germany
Heodes alciphron f. parallela Verity, 1946; Le Farfalle diurn. d'Italia 2: 42; TL: Italy
Heodes aciphron r. citragordius Verity, 1946; Le Farfalle diurn. d'Italia 2: 45; TL: Italy
denisei (Vigneau, 1949) (Heodes); Rev. franc. Lép. 12: 149; TL: France
rhaetica (Beuret, 1952) (Heodes); Mitt. ent. Ges. Basel (n.f.) 2: 86
cumanicus Szabó, 1956; Folia ent. hung. (n.s.) 9: 305, 340; TL: Hungary
burgundiae Bellenger & Descimon, 1977; Alexanor 10 (4): 165; TL: France
morvandica Bellenger & Descimon, 1977; Alexanor 10 (4): 166; TL: France
ochsenfeldinus (Betti, 1977) (Heodes); Alexanor 10 (2): 91; TL: West Germany

Chrysophanus alciphron ab. constricta; Schultz, 1905, Ent. Zs. 18 (34): 133
Heodes alciphron; [BARU], #347; [EBW]; [H&R]; [BOW]: pl. 9, f. 24; Yakovlev, 2012, Nota lepid. 35 (1): 67; [SZMN]
Lycaena (Heodes) alciphron; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25
Lycaena (Thersamolycaena) alciphron; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [RFEL]
Lycaena albescens; [CoL]
Lycaena alciphron aetnea; [CoL]
Lycaena alciphron bellieri; [CoL]
Lycaena alciphron burgundiae; [CoL]
Lycaena alciphron citragordius; [CoL]
Lycaena alciphron cumanicus; [CoL]
Lycaena alciphron denisei; [CoL]
Lycaena alciphron emarginata; [CoL]
Lycaena alciphron insignis; [CoL]
Lycaena alciphron mirabilis; [CoL]
Lycaena alciphron morvandica; [CoL]
Lycaena alciphron ochsenfeldinus; [CoL]
Lycaena alciphron paupercula; [CoL]
Lycaena alciphron rhaetica; [CoL]
Lycaena alciphron ruehli; [CoL]
Lycaena alciphron ultragordius; [CoL]
Lycaena alciphron veronicus; [iNaturalist]
Lycaena alciphron; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [MRS], 664; [Otakar Kudrna]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF] ; [Mario Maier]
Lycaena confluentiamultiplex; [CoL]
Thersamolycaena alciphron; [BRU2]: 126, pl. 58, f. 13-21

500x397 (~63Kb) underside Mt. Canigou, Pyrenees Orientales, France 27/7/96, Photo © Simon Coombes	533x400 (~57Kb) ♂ upperside A valley meadow at the Shadrikha Rivulet at the village Mel'nichikha, Novosibirsk District and Province, West Siberia, Russia. 17th May 1993, Photo © Oleg Kosterin	664x585 (~87Kb) ♂ upperside Russia, Moscow area, 11.06.2007, Photo © D. Smirnov
745x604 (~87Kb) ♂ Russia, Moscow area, 11.06.2007, Photo © D. Smirnov	784x684 (~112Kb) underside Russia, Moscow area, 9.6.2019 (36°25'E, 56°00'N), Photo © D. Smirnov

L. a. alciphron CEU, SEU, SW.Siberia, Altais, Sayan

?Chrysophanus alciphron madytus Fruhstorfer, 1917; Ent. Rundschau 34 (4): 17; TL: Sicily
?Chrysophanus alciphron veronius Fruhstorfer, 1917; Ent. Rundschau 34 (4): 17; TL: Pyrenees
?Chrysophanus alciphron epidelion Fruhstorfer, 1917; Ent. Rundschau 34 (4): 17; TL: Alpes Maritimes
?Chrysophanus alciphron deinareton Fruhstorfer, 1917; Ent. Rundschau 34 (4): 18; TL: Grans Sasso, Monti Sibillini

Lycaena alciphron alciphron; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]
Lycaena alciphron deinareton; [CoL]
Lycaena alciphron epidelion; [CoL]
Lycaena alciphron madytus; [CoL]
Lycaena alciphron veronius; [CoL]
Thersamolycaena alciphron alciphron; [BRU2], 126

L. a. gordius (Sulzer, 1776)

Papilio gordius Sulzer, 1776; Gesch. Ins. nach linn. Syst. (1): 146, (2): pl. 18, f. 7-8
Papilio columbanus de Prunner, 1798; Lepid. Pedemont.: 76
Polyommatus gordius ♂ ab. subtus-fere-radiata [= subtusfereradiata] Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 78
Polyommatus gordius ♀ ab. subtus-minus-punctata [= subtusminuspunctata] Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 79
Chrysophanus acliphron var. gordius ab. evanescens Gillmer, 1904; Soc. ent. 18 (23): 178; TL: Digne
Chrysophanus gordius ab. subfasciata Schultz, 1905; Ent. Zs. 18 (34): 133
Polyommatus gordius ab. diniensis Oberthür, 1906; Bull. Soc. ent. Fr. 1906: 26
Chrysophanus gordius ab. radiata Courvoisier, 1907; Zs. wiss. Insektenbiol. 3 (2): 37
Chrysophanus alciphron isokrates Fruhstorfer, 1909; Int. ent. Zs. 3 (21): 120; TL: Iselle, Simplon
Polyommatus gordius herrichii Oberthür, 1910; Ètud. Lépid. Comp. 4: 115
f. transversa Courvoisier, 1911; Ent. Zs. 24 (47): 255
sinestraradiata (Saussure, 1914) (Chrysophanus); Bull. Soc. Lép. Genève 3: 78
?Chrysophanus alciphron gordius sr. calabrus Verity, 1914; Boll. Soc. ent. ital. 45: 229
villaceus (Pionneau, 1926) (Chrysophanus); Echange 42: 7
magnifica (Pionneau, 1936) (Chrysophanus); Echange 52: 24 (repl. villaceus Pionneau, 1926)
subintermedia (Pionneau, 1937) (Chrysophanus); Echange 53: 11
occidentalis (Lalanne-Cassou, 1947) (Heodes); Rev. franc. Lep. 10: 24
phloeoides (Lalanne-Cassou, 1947) (Heodes); Rev. franc. Lep. 10: 27

Lycaena alciphron calabrus; [CoL]
Lycaena alciphron gordius; [CoL] ; [iNaturalist] ; [Heiner Ziegler] ; [Mario Maier]
Lycaena subtusfereradiata; [CoL]
Polyommatus gordius diniensis; Oberthür, 1910, Ètud. Lépid. Comp. 4: 667, pl. 38, f. 245

L. a. melibaeus (Staudinger, 1878) Caucasus, Transcaucasia

Polyommatus alciphron var. melibaeus Staudinger, 1878; Horae Soc. Ent. Ross. 14: 231
Chrysophanus alciphron fruginus Fruhstorfer, 1917; Ent. Rundschau 34 (4): 17; TL: Armenia
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75

Lycaena alciphron fruginus; [CoL] ; [iNaturalist]
Lycaena alciphron melibaeus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75; [iNaturalist]
Lycaena melibaeus; [CoL]
Thersamolycaena alciphron melibaeus; [BRU2], 126

L. a. granadensis (Ribbe, 1905) Spain

Chrysophanus gordius var. granadensis Ribbe, 1905; Soc. ent. 20 (18): 138; TL: Andalusia

Lycaena alciphron granadensis; [CoL]

L. a. heracleana (Blachier, 1908) Morocco (Toubkal Massif, Dj.Siroua)

Chrysophanus alciphron var. heracleana Blachier, 1908; Ann. Soc. ent. Fr. 77 (2): 217; TL: Atlas Mounains (Morocco)

Heodes alciphron heracleana; [BMAT]: 29, pl. 10, f. 12-15
Heodes alciphron heracleanus;
Lycaena alciphron heracleana; [CoL]

600x435 (~55Kb) upperside Morocco, Djebel Siroua, Anti-Atlas, Photo © Michel Tarrier

L. a. romanorum (Fruhstorfer, 1909) S.Italy

Chrysophanus alciphron romanorum Fruhstorfer, 1909; Int. ent. Zs. 3 (21): 120; TL: nr. Rome

Lycaena alciphron romanorum; [CoL]

L. a. gaudeolus (Fruhstorfer, 1909)

Chrysophanus alciphron gaudeolus Fruhstorfer, 1909; Int. ent. Zs. 3 (21): 120; TL: Tirol

Lycaena alciphron gaudeolus; [CoL] ; [iNaturalist]

L. a. naryna (Oberthür, 1910) N.Tian-Shan, Inner Tian-Shan, Dzhungarsky Alatau

Polyommatus gordius naryna Oberthür, 1910; Ètud. Lépid. Comp. 4: 115, 675, pl. 49, f. 407; TL: Fort Naryne, Turkestan

Lycaena alciphron naryna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [CoL]
Lycaena naryna; [CoL]
Thersamolycaena alciphron naryna; [BRU2], 126

L. a. chairemon (Fruhstorfer, 1917) Carpathians

Chrysophanus alciphron chairemon Fruhstorfer, 1917; Ent. Rundschau 34 (4): 16; TL: Herzegovina; Bulgaria, Orsowa

Lycaena alciphron chairemon; [CoL]
Thersamolycaena alciphron chairemon; [BRU2], 127

L. a. acutipennis Chou & Zhang, 1994 Xinjiang

Lycaena alciphron acutipennis Chou & Zhang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 771, 664, f. 73; TL: Xinjiang

L. a. trolli Korb, 2005 Transbaikalia

Lycaena alciphron trolli Korb, 2005; A catalogue of butterflies of the ex-USSR: 99
Heodes alciphron rubidus Korshunov, 1995; Butts. Asian Russia: 164 (preocc. Chrysophanus rubidus Behr, 1866)
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76 (nom. nud.); [RFEL]
Heodes alciphron rubidus; [BARU]

Heodes alciphron rubidus; [SZMN]
Lycaena alciphron trolli; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]
Thersamolycaena alciphron rubida; [BRU2], 126

Lycaena pavana (Kollar, [1844])

W.Himalayas, Kashmir - Kumaon, NW.India. See [maps] White-bordered Copper

Polyommatus pavana Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek 4: 416, pl. 5, f. 5-6

Lycaena (Rapsidia) pavana; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena pavana; [BIR], 301 (is not Nacaduba pavana Horsfield); [EBW]; [BOW]: pl. 178, f. 17
Lycaena pavana ?Horsfield;

Lycaena violaceus (Staudinger, 1892)

Altai, Sayan, Transbaikalia, Mongolia, N.China. See [maps]

Polyommatus dispar var. violaceus Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 315; TL: Kentei [Malakhanskiy mountain range, Kudara-Somon]

Lycaena (Thersamolycaena) violaceus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 75
Lycaena violacea; Bálint, 1996, Linneana Belg. 15 (6): 238; [iNaturalist]
Thersamolycaena violacea violacea; [BRU2], 126
Thersamolycaena violacea; [BRU2]: 126, pl. 58, f. 4-6
Thersamolycaena violaceus; [BARU], #345

533x400 (~42Kb) ♂ upperside A rocky southern slope of a piedmont hill of the mountain Gydyrgun on the northern bank of Lake Zun-Torei, Onon District, SE Chita Province (Dahuria), Transbaikalia, Siberia, Russia. 13th July 1996, Photo © Oleg Kosterin

Lycaena dabrerai Bálint, 1996 Mongolian Altai, Dzhungarian Gobi

Lycaena dabrerai Bálint, 1996; Linneana Belg. 15 (6): 237; TL: Mongolia, Bajanchongor aimak, Cagan Bogd ul, Tooroin bulag, 1500m

Thersamolycaena dabrerai; Yakovlev, 2012, Nota lepid. 35 (1): 67

Lycaena splendens (Staudinger, 1881)

Dzhungarsky Alatau, Tian-Shan. See [maps]

Polyommatus splendens Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 280; TL: Lepsa [Lepsy Valley near Cherkasskoe, Dzhungarsky Alatau, Kazakhstan]
Polyommatus splendens Alphéraky, 1882; Horae Soc. ent. Ross. 16 (3-4): 376, pl. 14, f. 12 (Staudinger i.l.)

Lycaena (Rapsidia) splendens; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena (Thersamolycaena) splendens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Lycaena splendens; [EBW]; [BOW]: pl. 206, f. 7; [iNaturalist]
Polyommatus splendens; [CoL]
Thersamolycaena splendens; [BRU2]: 126, pl. 58, f. 1-3

Lycaena kasyapa (Moore, 1865)

W.Himalayas, Kashmir, NW.India, Chitral - Mussoorie. See [maps] Green Copper

Chrysophanus kasyapa Moore, 1865; Proc. zool. Soc. Lond. 1865 (2): 506, pl. 31, f. 10

Lycaena (Rapsidia) kasyapa; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Bálint, 1992, Linn. Belg. 13 (8): 400
Lycaena kasyapa; [BIR], 301; [EBW]; [BOW]: pl. 178, f. 16; Tshikolovets, 2017, Zootaxa 4358 (1): 115; [CoL] ; [iNaturalist]

Lycaena aeolus Wyatt, 1961

W.Pmairs (Shugnansky, Ishkashimsky Mts.), Afghanistan. See [maps]

Lycaena aeolus Wyatt, 1961; J. Lep. Soc. 15 (1): 17; TL: Bala-Quran, Anjuman Valley, Hindu Kush Mts., 14300ft, Afghanistan

Lycaena (Thersamolycaena) aeolus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Lycaena aeolus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name); [CoL]
Thersamolycaena aeolus; [BRU2]: 127, pl. 57, f. 40-42
Thersamonia aeolus;

L. a. aeolus

Lycaena aeolus aeolus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

L. a. aeolides (Churkin, 1999) Ghissar

Thersamolycaena aeolides Churkin, 1999; Atalanta 29: 125, pl. IVb, f. 1; TL: Ghissar, Fanskie Mts.

Lycaena aeolus aeolides; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Thersamolycaena aeolides; [BRU2]: 127, pl. 57, f. 43-45

Lycaena standfussi (Grum-Grshimailo, 1891)

Tibet - W.China. See [maps]

Polyommatus standfussi Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 450

Lycaena (Rapsidia) standfussi; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name)
Lycaena standfussi; [BOW]: pl. 206, f. 8; [CoL] ; [iNaturalist]

1300x984 (~256Kb) ♀ upperside China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

900x749 (~122Kb) ♂ underside China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

L. s. subbrunnea (South, 1913)

Chrysophanus subbrunnea South, 1913; J. Bomb. nat. Hist. Soc. 22 (3): 607

Lycaena standfussi subbrunnea; Huang, 2001; [CoL]

L. s. sinensis (Nordström, 1935)

standfussi sinensis (Nordström, 1935) (Chrysophanus); Ark. Zool. 27A (7): 3

Lycaena standfussi sinensis; [CoL]

Lycaena adbayar (Churkin, 2004)

Thersamolycaena adbayar Churkin, 2004; Helios 5: (75-90)

Thersamolycaena adbayar; Yakovlev, 2012, Nota lepid. 35 (1): 67

Lycaena otbayar (Churkin, 2004)

Mongolia. See [maps]

Thersamolycaena otbayar Churkin, 2004; Helios 5: (75-90); TL: Mongolia, Gobi-Altaisky aimak, Gobi-Altai mts, 30km S Biger village, 2700-3000m

Subgenus Heodes Dalman, 1816

Heodes Dalman, 1816; K. VetenskAcad. Handl. 1816 (1): 63; TS: Papilio virgaureae Linnaeus
- = Lycaena; Stempffer, 1967, Bull. Br. Mus. nat. Hist. (Ent) Suppl. 10: 263; [NL4A], 137
Chysoptera Zincken, 1817; Allgem. Literaturzeitung, 1817: 75; TS: Papilio virgaureae Linnaeus
- = Heodes; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
- = Subgenus Heodes (chrysoptera); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
- = Lycaena; [NL4A], 137
Loweia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 314; TS: Papilio dorilis Hufnagel
- = Subgenus Heodes (chrysoptera); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
- = Lycaena; [NL4A], 138
- = Subgenus Heodes; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Palaeoloweia Verity, 1934; Ent. Rec. J. Var. (Suppl) 46: 13 (repl. Loweia Tutt, 1906); TS: Papilio dorilis Hufnagel
- = Loweia; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
- = Subgenus Heodes (chrysoptera); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
- = Lycaena; [NL4A], 138
Palaeochrysophanus Verity, 1943; Le Farfalle diurn. d'Italia 2: 23, 64; TS: Papilio hippothoe Linnaeus
- = Lycaena; [NL4A], 138
- = Subgenus Heodes; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Palaeochrysophanus Verity, 1934; Ent. Rec. J. Var. (Suppl): (13) nota [invalid]

Subgenus Heodes (Lycaena); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Chrysoptera; Tutt, 1906, (missp.); [NHM-BMW] ; [NHM-BMW]
Chysoptera; [NHM-BMW]
Heodes; [NHM-BMW]
Heodes (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
Loweia; [CoL] ; [NHM-BMW]
Loweia (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
Lycaena (Heodes); [P&N23]
Palaeochrysophanus; [CoL] ; [NHM-BMW] ; [NHM-BMW]
Palaeochrysophanus (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Palaeoloweia; [NHM-BMW]

Lycaena virgaureae (Linnaeus, 1758)

EU, Siberia, Asia Minor, C.Asia, Mongolia. See [maps] Loistokultasiipi Manto de Oro Argus Satiné Vitfläckig guldvinge Dukatenfalter Scarce Copper Guldvinge Großer Feuerfalter Czerwończyk dukacik

Papilio virgaureae Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 484; TL: Sweden
lunulata (Courvoisier, 1903) (Chrysophanus); Mitt. schweiz. ent. Ges. 11: 23
voelschowi Gillmer, 1903²; Arch. Ver. Nat. Mecklenburg: 122
Chrysophanus virgaureae ♂ ab. guttata Schultz, 1904; Ent. Zs. 18 (24): 93
Chrysophanus virgaureaea ab. chryzon Oberthür, 1905; Bull. Soc. ent. Fr. 1905: 56; TL: Frankfurt
Chrysophanus virgaureae ab. virginalis Oberthür, 1905; Bull. Soc. ent. Fr. 1905: 56; TL: Silesia
Chrysophanus virgaureae ab. elongata Schultz, 1905; Ent. Zs. 18 (33): 130 (Courv.)
Chrysophanus virgaureae ♂ ab. sincera Schultz, 1905; Ent. Zs. 18 (33): 131
Chrysophanus virgaureae ♀ ab. milena Schultz, 1905; Ent. Zs. 18 (33): 131
Chrysophanus virgaureae ♀ ab. lateradiata Schultz, 1905; Ent. Zs. 18 (33): 131
parallela (Courvoisier, 1907) (Chrysophanus); Zs. wiss. Insektenbiol. 3 (2): 36
Heodes virgaureae var. suldensis Tutt, 1909; Ent. Rec. J. Var. 21: 229
punctifera (Courvoisier, 1911) (Chrysophanus); Ent. Zs. 24 (42): 233
paucipuncta (Courvoisier, 1911) (Chrysophanus); Ent. Zs. 24 (42): 236
pleuripuncta (Courvoisier, 1911) (Chrysophanus); Ent. Zs. 24 (42): 236
Chrysophanus virgaureae f. tripuncta Closs, 1920; Int. ent. Zs. 14 (16): 122; TL: Tegel
Chrysophanus virgaurea[sic] ab. bergeri Schindler, 1922; Ent. Anz. 2 (5): 57
emilianus (Turati, 1923) (Chrysophanus); Atti Soc. ital. Sci. nat. 62: 42; TL: Apennines
quercii (Turati, 1923) (Chrysophanus); Atti Soc. ital. Sci. nat. 62: 42; TL: Apennines
alba (Schoenfeld, 1924) (Chrysophanus); Int. ent. Zs. 18: 40; TL: Bavaria
subtusrufomarginata Graves & Hemming, 1928; Entomologist 61 (777): 25
subtuorufomarginata Graves & Hemming, 1928; Entomologist 61 (777): 31
vera Graves & Hemming, 1928; Entomologist 61 (778): 61
pyrenaeicola Graves & Hemming, 1928; Entomologist 61 (780): 104; TL: N.Pyrenees
Heodes virgaureae gravesi Verity, 1929; Bull. Soc. ent. Fr. 34 (7): 129
Heodes virgaureae gravesica Verity, 1929; Bull. Soc. ent. Fr. 34 (7): 129; TL: Lozere
Heodes virgaureae mediomontana Verity, 1929; Bull. Soc. ent. Fr. 34 (7): 131; TL: Basses-Alpes
Heodes virgaureae pyrenemontana Verity, 1929; Bull. Soc. ent. Fr. 34 (7): 131; TL: Pyrenees Orientales
delicata (Higgins, 1930) (Chrysophanus); Entomologist 63: 99; TL: Piedmont
multipunctata (Warnecke, 1942) (Chrysophanus); Dt. ent. Z. Iris 56: 103
rhehana (Heydemann, 1953) (Chrysophanus); Dt. ent. Z. Iris 55: 99
costipuncta (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 11
pseudomiegi (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 11
quadrilunulata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 11
suprabasielongata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 12
supracentroelongata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 12
ultraseriata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 12
subtusdiscoelongata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 13

Chrysophanus virgaureae; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 111; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 31
Heodes virgaureae; [BARU], #348; Yakovlev, 2012, Nota lepid. 35 (1): 68; [H&R]; [BOW]: pl. 9, f. 26, 28, 29; [BRU2]: 124, pl. 58, f. 25-36
Lycaena (Heodes) virgaureae; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]
Lycaena bergeri; [CoL]
Lycaena subtuorufomarginata; [CoL]
Lycaena subtusdiscoelongata; [CoL]
Lycaena suldensis; [CoL]
Lycaena virgaureae chrysorrhoa; [CoL]
Lycaena virgaureae costipuncta; [CoL]
Lycaena virgaureae delicata; [CoL]
Lycaena virgaureae emilianus; [CoL]
Lycaena virgaureae gravesi; [CoL]
Lycaena virgaureae gravesica; [CoL]
Lycaena virgaureae mediomontana; [CoL]
Lycaena virgaureae multipunctata; [CoL]
Lycaena virgaureae pseudomiegi; [CoL]
Lycaena virgaureae pyrenaeicola; [CoL]
Lycaena virgaureae pyrenemontana; [CoL]
Lycaena virgaureae quadrilunulata; [CoL]
Lycaena virgaureae quercii; [CoL]
Lycaena virgaureae rhenana; [CoL]
Lycaena virgaureae subtusdiscoelongata; [CoL]
Lycaena virgaureae suprabasielongata; [CoL]
Lycaena virgaureae supracentroelongata; [CoL]
Lycaena virgaureae tripuncta; [CoL]
Lycaena virgaureae ultraseriata; [CoL]
Lycaena virgaureae vera; [CoL]
Lycaena virgaureae; [SPRK]; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [MRS], 664; [Otakar Kudrna]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF] ; [Mario Maier]
Polyommatus virgaureae virginalis; Oberthür, 1910, Ètud. Lépid. Comp. 4: 135, 667, pl. 37, f. 244

450x327 (~42Kb) ♂ upperside Mt. Canigou, Pyrenees Orientales, France 27/07/96, Photo © Simon Coombes	680x680 (~60Kb) ♀ upperside FINLAND: Ab: Suomusjärvi, 669:31, 27.7.1997, Photo © Markku Savela	600x436 (~25Kb) ♂ upperside FINLAND: Kb: Kiihtelysvaara, 692:68, 11.7.1997, Photo © Markku Savela
600x688 (~52Kb) ♂ underside FINLAND: Kb: Kiihtelysvaara, 692:68, 11.7.1997, Photo © Markku Savela	1400x968 (~124Kb) ♀ upperside FINLAND, Ab: Suomusjärvi, 669:31, 2.8.1998, Photo © Markku Savela	600x803 (~46Kb) ♂ upperside Finland, Sipoo 670:39, 12.7.1979, Photo © Pertti Pakkanen
596x400 (~42Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	596x400 (~53Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch	979x734 (~91Kb) ♂ Lithuania: Dzukija, Marcinkonys-Zervynos, 15.7.2008 (on Thymus pulegioides), Photo © Harri Arkkio
745x938 (~80Kb) ♂ upperside Lithuania: Dzukija, Marcinkonys, railwaystation, 16.7.2008, Photo © Harri Arkkio	914x685 (~73Kb) ♂ underside Lithuania: Dzukija, Marcinkonys-Zervynos, 15.7.2008, Photo © Harri Arkkio	680x610 (~96Kb) ♂ underside Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
877x1080 (~196Kb) ♀ Russia, Khabarovskiy Kray, Khabarovsk southern env., Korfovskiy settlement. 11th July 2014, Photo © Oleg Kosterin	886x865 (~258Kb) ♀ upperside Russia, Khabarovskiy Kray, Khabarovsk southern env., Korfovskiy settlement. 11th July 2014, Photo © Oleg Kosterin	1016x1168 (~261Kb) ♀ underside Russia, Khabarovskiy Kray, Khabarovsk southern env., Korfovskiy settlement. 11th July 2014, Photo © Oleg Kosterin
500x444 (~23Kb) ♀ FINLAND: Ab: Suomusjärvi, 669:31, 18.8.1996, Photo © Markku Savela	500x389 (~22Kb) ♂ upperside Finland, Sotkamo, Pihlajapuro, 7/1996, Photo © Tero Piirainen	299x299 (~11Kb)

L. v. virgaureae EU, W.Siberia, Caucasus, Tarbagatai, Altai

Lycaena virgaureae oranula Freyer, 1845; Neuere Beitr. Schmett. 5: pl. 455, f. 1-2
Polyommatus estonica Huene, 1883; Ent. Nachr. 9 (4): 49
?Polyommatus virgaureae var. estonica ab. ♂ apicepunctata Huene, 1901; Stettin Ent. Ztg 62 (1-6): 154
Polyommatus virgaureae var. estonica ab. ♀ albopunctata Huene, 1901; Stettin Ent. Ztg 62 (1-6): 154
Chrysophanus virgaureae inalpinus Verity, 1913; J. Linn. Soc. Zool. Lond. 32 (215): 183
Lycaena (Heodes) virgaureae balcanicola Graves, 1928; Entomologist 61 (780): 106; TL: S.Bulgaria, Kostenetz, 3000ft
Lycaena virgaureae pyronitens Szabó, 1956; Folia ent. hung. (n.s.) 9: 299, 340; TL: Hungary
- = Lycaena virgaureae balcanicola; Bálint, 1991, Janus Pannonius Múz. 35: 49

Chrysophanus virgaureae inalpinus; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 40
Heodes virgaureae virgaureae; [BRU2], 124
Lycaena albopunctata; [CoL]
Lycaena virgaureae oranula; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 32 (name)
Lycaena virgaureae pyronitens; [CoL]
Lycaena virgaureae virgaureae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Polyommatus virgaureae oranula; Oberthür, 1910, Ètud. Lépid. Comp. 4: 135
balcanicola; Abadjiev, 2002, Neue Ent. Nachr. 53: 18

L. v. montana (Meyer-Dür, 1852)

Polyommatus virgaureae var. montana Meyer-Dür, 1852; Verz. Schmett. Schweiz 1: 53

Chrysophanus virgaureae montana; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 111
Lycaena virgaureae montana; [CoL]

L. v. miegii (Vogel, 1857) Spain

miegii (Vogel, 1857) (Polyommatus); Zeit. Ges. Isis: 201, pl. 6; TL: Spain

Chrysophanus virgaureae miegi; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 42
Lycaena (Heodes) miegi; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25
Lycaena virgaureae miegii; [CoL]
Polyommatus virgaureae miegii; Oberthür, 1910, Ètud. Lépid. Comp. 4: 135

L. v. zermattensis (Fallou, 1865)

virgaureae zermattensis (Fallou, 1865) (Polyommatus); Ann. Soc. Ent. Fr. (4) 5: 101, pl. 2, f. 3
Chrysophanus virgaureae var. zermattensis ab. caeruleopunctata Schultz, 1905; Ent. Zs. 18 (33): 131
Polyommatus virgaureae zermattensis ♀ ab. cuneifera Oberthür, 1910; Ètud. Lépid. Comp. 4: 135
Chrysophanus virgaureae zermattensis f. seriata Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 194
Chrysophanus virgaureae zermattensis f. fredegunda Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 194
Chrysophanus virgaureae zermattensis f. onka Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 195

Chrysophanus virgaureae fredegunde[sic, fredegunda]; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 112
Chrysophanus virgaureae seriata; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 112
Chrysophanus virgaureae zermattensis; Fruhstorfer, 1908, Int. ent. Zs. 2 (29): 194; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 112
Lycaena fredegunda; [CoL]
Lycaena virgaureae onka; [CoL]
Lycaena virgaureae seriata; [CoL]
Lycaena virgaureae zermattensis; [CoL]

L. v. apennina (Calberla, 1888)

Polyommatus virgaureae var. apennina Calberla, 1888; Corr.-Bl. Ent. Ver. Iris 1 (1) (4): 125; TL: Apennin

Crysophanus virgaureae apennina; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 42
Lycaena virgaureae apennina; [CoL]

L. v. virgaureola (Staudinger, 1892) Sayan, Transbaikalia, Amur, Ussuri

Polyommatus virgaureae virgaureola Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 314; TL: Kentei?
Heodes steni Bryk, 1946
Chrysophanus virgaureae mongolica Kurenzov, 1970

Chrysophanus virgaureae virgaureola; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 42
Heodes virgaureae virgaureola; [BARU](note) , ; [BRU2], 124
Lycaena virgaureae virgaureola; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]; [CoL]

400x300 (~32Kb) ♂ ♀ underside A forb meadow on a SE slope 12 km SE of the village Gazimurskii Zavod, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 24th July 1997, Photo © Oleg Kosterin

400x451 (~32Kb) underside A meadow, with Medicago dominating, on the Tes-Khem River right bank in front of the Shivilig-Khem valley, the Ubsu-Nur intermontane hollow, Tes-Khem District, Tuva Republic, Siberia, Russia. 9th July 1990, Photo © Oleg Kosterin

L. v. aureomicans (Heyne, 1897)

Polyommatus virgaureae var. aureomicans Heyne, 1897; Soc. Ent. 12 (2): 9

Lycaena aurcomicaus; [CoL]
Lycaena virgaureae aureomicans; [CoL]
Polyommatus virgaureae aureomicans; Oberthür, 1910, Ètud. Lépid. Comp. 4: 135

L. v. armeniaca (Bang-Haas, 1906) Armenia (highlands)

Chrysophanus virgaureae armeniaca Bang-Haas, 1906; Dt. ent. Z. Iris, 19 (3): 128

Heodes virgaureae armeniaca; [BRU2], 124
Lycaena virgaureae armeniaca; [CoL]

L. v. athanagild (Fruhstorfer, 1908) Engadin

Chrysophanus virgaureae athanagild Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 194; TL: Engadin

Chrysophanus virgaureae athanagild; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 112; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 38
Lycaena virgaureae athanagild; [CoL]

L. v. juvara (Fruhstorfer, 1908) Bayern

Chrysophanus virgaureae juvara Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 195; TL: Bayern, Passau
Chrysophanus virgaureae juvara f. galsnintha Fruhstorfer, 1908; Int. ent. Zs. 2 (29): 195; TL: S.Tirol, Vallombroso, Italy

Chrysophanus virgaureae juvara; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 33-34
Lycaena virgaureae galsnintha; [CoL]
Lycaena virgaureae juvara; [CoL]

L. v. osthelderi (Fruhstorfer, 1909)

Chrysophanus virgaureae osthelderi Fruhstorfer, 1909; Int. ent. Zs. 3 (20): 113; TL: onazzatal, Tosafall

Chrysophanus virgaureae osthelderi; Vorbrodt & Müller-Rutz, 1911, Die Schmett. Schweiz 1: 112; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 39
Lycaena virgaureae osthelderi; [CoL]

L. v. alexandrae (Fruhstorfer, 1909) Ural

Chrysophanus virgaureae alexandrae Fruhstorfer, 1909; Int. ent. Zs. 3 (21): 120; TL: Ural, Turgojak
- = Lycaena virgaureae virgaureae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

Chrysophanus virgaureae alexandrae; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 42
Heodes virgaureae alexandrae; [BARU](note) ,

L. v. pelusiota (Fruhstorfer, 1910)

Chrysophanus virgaureae pelusiota Fruhstorfer, 1910; Ent. Zs. 24 (26): 144; TL: Cogne

Chrysophanus virgaureae pelusiota; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 39
Lycaena virgaureae pelusiota; [CoL]

L. v. chrysorhoas (Fruhstorfer, 1917)

Chrysophanus virgaureae chrysorhoas Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 33

L. v. cissites (Fruhstorfer, 1917)

Chrysophanus virgaureae cissites Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 34

Lycaena virgaureae cissites; [CoL]

L. v. theages (Fruhstorfer, 1917)

Chrysophanus virgaureae theages Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 40

Lycaena theages; [CoL] ; [iNaturalist]
Lycaena virgaureae theages; [CoL]

L. v. lena (Kurenzov, 1970) C.Urals, C.Siberia, Far East

Chrysophanus virgaureae lena Kurenzov, 1970; ; TL: C. & E. Yakutia

Heodes virgaureae lena; [BARU](note) , ; [BRU2], 124
Lycaena virgaureae lena; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

L. v. denizae Eckweiler & Rose, 1993 SW.Turkey

Lycaena (Heodes) virgaureae denizae Eckweiler & Rose, 1993; Nachr. Ent. Ver. Apollo NF 13 (3a): 356; TL: Turkey, Isparta, Egridir, Aksu, Dedegöl Dag

L. v. mikaeli Tsvetkov, 2010 Caucasus

Lycaena virgaureae mikaeli Tsvetkov, 2010; Caucasian Ent. Bull. 6 (1): 99 (repl. Chrysophanus virgaureae caucasica Jachontov, 1908)
Chrysophanus virgaureae caucasica Jachontov, 1908; Revue Russe Ent. 8: 291 (preocc. Lycaena coridon caucasica Lederer, 1870)
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

Heodes virgaureae caucasica; Koçak, 1977, Atalanta 8 (1): 46
Lycaena virgaureae mikaeli; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

Lycaena tityrus (Poda, 1761)

SEU, CEU, S.Russia, Transcaucasus, Altai. See [maps] Sooty Copper Manto oscuro Bienenflater Czerwończyk uroczek Dansk Guldvinge

Papilio tityrus Poda, 1761; Ins. Mus. Graecensis: 77; TL: Austria, Graz
acrion Pontoppidan, 1763
Papilio alexis Scopoli, 1763; Ent. Carniolica: 179
Papilio dorilis Hufnagel, 1766; Berlin. Mag. 2: 68
- = ; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
dorilas Rottemburg, 1775
Papilio circe Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 181
Papilio xanthe Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 181
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 495
Papilio phocas Rottemburg, 1775; Der Naturforscher 6: 29
Papilio rubi Fourcroy, 1785; Ent. Paris. 2: 245
Papilio garbus Fabricius, 1787; Mantissa Insectorum 2: 81
Polyommatus xanthe var. canidia Gerhard, 1850; Versuch Mon. europ. Schmett. (3): 7, pl. 10, f. 2
- = tityrus; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 91
Polyommatus circe var. subalpina Speyer, 1851; Stettin Ent. Ztg 12: 339
- = (subalpinus); Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 441
Polyommatus xanthe var. Bellier, 1859; Ann. Soc. ent. Fr. (3) 7: 188, pl. 5 II, f. 3; TL: Basses-Alpes
?hypoxanthe (Kirby, 1862) (Chrysophanus); Man. Eur. Butt.: 91, f. 11; TL: ?Ukraine
- = Lycaena dorilis; Kirby, 1882, Eur. Butts Moths: 54
Chrysophanus dorilis var. locarmensis Tutt, 1903; Ent. Rec. J. Var. 15: 224; TL: Locarno
Chrysophanus dorilis ab. striata Gillmer, 1904; Soc. ent. 18 (23): 179
Chrysophanus dorilis f. upoleuca Verity, 1904; Entomologist 37 (490): 58
Polyommatus dorilis var. brantsi Ter Haar, 1900; Tijdschr. Ent. 43: 237, pl. 14, f. 2
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 77
Polyommatus dorilis var. uyeni Ter Haar, 1900; Tijdschr. Ent. 43: 237, pl. 14, f. 3
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 480
Chrysophanus dorilis ab. strandi Schultz, 1903; Nyt mag. naturv. 41: 25
Chrysophanus dorilis ab. ♂ fulvomarginalis Schultz, 1905; Ent. Zs. 18 (34): 134
Chrysophanus dorilis ab. ♀ fusca Gillmer, 1908; Ent. Wochenbl. 25 (5): 20
striata (Ksienschopolski, 1911) (Chrysophanus); Trudy obshch.Izsl. Volyni Zitomir 8: 56
straminea (Blachier, 1912) (Chrysophanus); Bull. Soc. Lep. Genève 2: 51
Loweia dorilis r. italorum Verity, 1919; Ent. Rec. J. Var. 31: 29
Chrysophanus dorilis reverdini Stauder, 1921; Dt. ent. Z. Iris 35 (1/2): 30; TL: "Faito und Aspromonte, 400-950m"
- = ; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 400
aricia (Cabeau, 1922) (Chrysophanus); Rev. mens. Soc. ent. Namur. 1922: 3
Loweia dorilis italorum f. italaveris Verity, 1924; Ent. Rec. J. Var. 36: 108
argentea (Gussich, 1930) (Chrysophanus); Glasnik. hrv. prirodosl. dr. Zagreb 29: 222; TL: Croatia
semiobscurior (Pionneau, 1931) (Chrysophanus); L'Echange 47: 7
intermedia (Pionneau, 1931) (Chrysophanus); L'Echange 47: 7
Palaeoloweia tityrus pallidepicta Verity, 1934; Ent. Rec. J. Var. (Suppl) 46: 16
Palaeoloweia tityrus praebleusei Verity, 1934; Ent. Rec. J. Var. (Suppl) 46: 17; TL: Rhone valley
Chrysophanus dorilis ab. splendens Wize, 1936; Polskie Pismo Ent. 13: 107
Heodes tityrus ab. intermedia Lempke, 1936; Tijdschr. Ent. 79: 295 (preocc.)
Heodes tityrus ab. flavescens Lempke, 1937; Tijdschr. Ent. 80: 303 (repl. Heodes tityrus ab. intermedia Lempke, 1936)
Palaeoloweia opisthochros Verity, 1939; Lambillionea 39: 186; TL: Persia
?infracana (Verity, 1946) (Heodes); Le Farfalle diurn. d'Italia 2: 34
?infraflava (Verity, 1946) (Heodes); Le Farfalle diurn. d'Italia 2: 34
Heodes tityrus r. mixtalpina Verity, 1946; Le Farfalle diurn. d'Italia 2: 39
ab. cuneifera (Popescu-Gorj, 1948) (Chrysophanus); Notat. biol. Bucharest 6: 142; TL: Rumania
- = (subalpinus); Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 441
Chrysophanus dorilis ab. vilmae Komárek, 1949; Acta Soc. ent. Cal. 46: 149
fulvosignata (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 36
postlocarnensis (Beuret, 1954) (Heodes); Lycaeniden Schweiz 1: 41
Heodes (Chrysophanus) tiryrus f. centroelongata Lempke, 1954; Tijdschr. Ent. 97: 323
Heodes (Chrysophanus) tiryrus f. arcuata Lempke, 1954; Tijdschr. Ent. 97: 323
Heodes (Chrysophanus) tiryrus f. costojuncta Lempke, 1954; Tijdschr. Ent. 97: 324
Heodes (Chrysophanus) tiryrus f. limbojuncta Lempke, 1954; Tijdschr. Ent. 97: 324
Heodes (Chrysophanus) tiryrus f. imostriata Lempke, 1954; Tijdschr. Ent. 97: 324
Heodes (Chrysophanus) tiryrus f. basinovopuncta Lempke, 1954; Tijdschr. Ent. 97: 324
Heodes (Chrysophanus) tiryrus f. parvipuncta Lempke, 1954; Tijdschr. Ent. 97: 324
Heodes (Chrysophanus) tiryrus f. remota Lempke, 1954; Tijdschr. Ent. 97: 325

Chrysophanus dorilis reverdini; Stauder, 1924, Soc. Ent. 39 (6): 24
Heodes tityrus; [H&R]; [BARU], #350; [EBW]; [BOW]: pl. 9, f. 25, 27; [BRU2]: 124, pl. 60, f. 1-3
Loweia dorilis brantsi; [CoL]
Loweia dorilis intermedia; [CoL]
Loweia dorilis italaveris; [CoL]
Loweia dorilis italorum; [CoL]
Loweia dorilis italoveris; [CoL]
Loweia dorilis reverdini; [CoL]
Loweia dorilis subalpina; [CoL]
Loweia dorilis; [CoL]
Loweia tityrus arcuata; [CoL]
Loweia tityrus basinovopuncta; [CoL]
Loweia tityrus centroelongata; [CoL]
Loweia tityrus costojuncta; [CoL]
Loweia tityrus fulvosignata; [CoL]
Loweia tityrus imostriata; [CoL]
Loweia tityrus infracana; [CoL]
Loweia tityrus infraflava; [CoL]
Loweia tityrus limbojuncta; [CoL]
Loweia tityrus mixtalpina; [CoL]
Loweia tityrus opisthochros; [CoL]
Loweia tityrus pallidepicta; [CoL]
Loweia tityrus parvipuncta; [CoL]
Loweia tityrus praebleusei; [CoL]
Loweia tityrus remota; [CoL]
Loweia tityrus; [CoL]
Lycaena (Heodes) subalpina; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25
Lycaena (Heodes) tityrus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [P&N23]
Lycaena hypoxanthe; [CoL]
Lycaena subalpina; [MBE]
Lycaena tityrus dorilis; [iNaturalist]
Lycaena tityrus italorum; [iNaturalist]
Lycaena tityrus locarnensis; [iNaturalist]
Lycaena tityrus subalpina; [iNaturalist]
Lycaena tityrus; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF] ; [Mario Maier]
Papilio xanthe; Fabricius, 1787, Mantissa Insectorum 2: 81
Polyommatus dorilis; Staudinger, 1861, in Staudinger & Wocke, Cat. Lep. (ed. 1): 4
Polyommatus uyeni; [CoL]
?Papilio circe; Hübner, [1799-1800], Samml. eur. Schmett. [1]: 55, pl. 67, f. 334-336

Misidentified/misapplied:

Papilio xanthe; Lang, 1789, Verz. Schmett. Gegend. Augsburg (ed. 2): 52
- = Helleia helle; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 495

443x400 (~30Kb) ♀ underside The floodland on the Om' River right bank in the Omsk City suburbs, West Siberia, Russia. 14th August 1995, Photo © Oleg Kosterin	870x597 (~44Kb) ♂ upperside Paetz, Berlin, Germany, 7-90, Photo © S. Shuichi Haupt	906x806 (~156Kb) upperside Russia, North Caucasus, Adygea Republic, Maikop, Khanskaya stanitsa, Khanskie ponds. 14th July 2008, Photo © Oleg Kosterin
570x570 (~78Kb) upperside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov

L. t. tityrus

Lycaena tityrus tityrus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

L. t. orientalis (Staudinger, 1881) Ala Tau

Polyommatus dorilis orientalis Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 281

Chrysophanus dorilis v. orientalis; Staudinger, 1901, Cat. Lep. palaearct. Faunengeb. 1: 75
Loweia dorilis orientalis; [CoL]
Lycaena tityrus orientalis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [iNaturalist]

L. t. argentifex Bálint, 1990

Lycaena (Heodes) tityrus argentifex Bálint, 1990; Janus Pannonius Múz. 34: 47, 55; TL: "Békási-szoros, Kupás-p."

Lycaena tityrus argentifex; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76

Lycaena bleusei (Oberthür, 1884)

E.Spain. See [maps]

Polyommatus bleusei Oberthür, 1884; Étud. d'Ent. 8: 15; TL: Spain

Loweia dorilis bleusei; [CoL]
Lycaena (Heodes) bleusei; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25
Lycaena bleusei; Martins, 2011, Thesis: 33; [iNaturalist] ; [LEPIFORUM] ; [MBE]

Lycaena ottomanus (Lefèbvre, 1830)

gr, Asia Minor, Balkans. See [maps] Grecian Copper

Polyommatus ottomanus Lefèbvre, 1830; Rev.?/Mag. Zool. 1830: pl. 19
legeri Freyer, 1830; Beitr. eur. Schmett. 3 (17-24): pl. 133, f. 1

Heodes ottomanus; [EBW], (name); [H&R]
Lycaena (Heodes) ottomana; Cassulo, Mensi & Balletto, 1989, Nota Lepid. Suppl. 1: 25
Lycaena (Heodes) ottomanus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena ottomana; [Otakar Kudrna]; [MBE]
Lycaena ottomanus; [iNaturalist] ; [LEPIFORUM]
Polyommatus ottomanus; [CoL]

The palaeochrysophanus species-group

Lycaena hippothoe (Linnaeus, 1761)

EU, Russia, Siberia, Amur. See [maps] Ketokultasiipi Kosmosvinge Manto de Cobre Czerwończyk płomieniec Violettkantad guldvinge Purple-edged Copper

Papilio hippothoe Linnaeus, 1761; Fauna Suecica (Edn 2): 274; TL: Sweden
Polyommatus hippothoe var. groningana Ter Haar, 1900; Tijdschr. Ent. 43: 242
Polyommatus hippothoe ab. ♂ radiata Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 69
Chrysophanus hippothoe ab. argenteola Schultz, 1903; Nyt mag. naturv. 41: 23
Chrysophanus hippothoe ab. decurtata Schultz, 1903; Nyt mag. naturv. 41: 24
Chrysophanus hippothoe ab. ornata Züllich, 1929; Zs. Öst. EntVer. 14 (5): 51; TL: Rilogebirge, 1400m
Heodes hippothoe ab. parvipuncta Lempke, 1936; Tijdschr. Ent. 79: 292
Palaeochrysophanus hippothoe f. infragrisea Lempke, 1954; Tijdschr. Ent. 97: 340
Palaeochrysophanus hippothoe f. denigrata Lempke, 1954; Tijdschr. Ent. 97: 340

Heodes hippothoe; [BARU], #349; [BRU2]: 125, pl. 60, f. 5-12; Yakovlev, 2012, Nota lepid. 35 (1): 67; [SZMN]
Lycaena (Heodes) hippothoe; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]
Lycaena hippothoe italica; [iNaturalist]
Lycaena hippothoe; [SPRK]; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF] ; [Mario Maier]
Palaeochrysophanus hippothoe denigrata; [CoL]
Palaeochrysophanus hippothoe infragrisea; [CoL]
Palaeochrysophanus hippothoe; [H&R], 249; [EBW]; [BOW]: pl. 10, f. 9, 10; [MRS], 664; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [CoL]
Paleochrysophanus[sic] hippothoe; [BMAT]: 102, pl. 32, f. 11 (not in N.Africa)

500x455 (~31Kb) ♀ upperside Finland, Tampere, Rusko, 6819:335, 12.7.1996, Photo © Tero Piirainen	400x359 (~21Kb) ♂ upperside Finland, Tampere, Rusko, 6819:335, 12.7.1996, Photo © Tero Piirainen	550x588 (~35Kb) ♂ upperside FINLAND: Kl: Parikkala, 683:64, 1.7.1997, Photo © Markku Savela
500x564 (~35Kb) ♂ underside FINLAND: Kl: Parikkala, 683:64, 1.7.1997, Photo © Markku Savela	600x742 (~40Kb) ♀ upperside Finland, Sipoo 670:39, 26.6.1980, Photo © Pertti Pakkanen	600x719 (~49Kb) ♂ upperside Finland, Sipoo 670:40, 20.6.1980, Photo © Pertti Pakkanen
1200x834 (~104Kb) underside Finland, Sipoo 670:39, 24.6.1981, Photo © Pertti Pakkanen	694x660 (~93Kb) ♀ Russia, Moscow area, 16.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov	1119x540 (~90Kb) ♂ Russia, Moscow area, 17.06.2007, Photo © D. Smirnov

L. h. hippothoe EU, W.Siberia, Dzhungarsky Alatau, Tarbagatai

Papilio chryseis Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 181
Papilio euridice Rottemburg, 1775; Der Naturforscher 6: 28, n. 17
Polyommatus chryseis var. confluens Gerhard, 1850; Versuch Mon. europ. Schmett. (2): 5, pl. 8, f. 1a-d
Polyommatus chryseis ♂ ab. fere-confluens Oberthür, 1896; Étud. d'Ent. 20: pl. 5, f. 81
alciphronides Krulikovsky, 1897
Chrysophanus hippothoe ab. spadona Krulikovsky, 1909; Rev. russ. ent. 9 (3): 300
Polyommatus chryseis ab. subtusimpunctata Oberthür, 1910; Ètud. Lépid. Comp. 4: 129
- = hippothoe; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 444
bourdouxhei (Cabeau, 1924) (Chrysophanus); Rev. mens. Soc. ent. Namur. 24: 18

Heodes hippothoe hippothoe; [BRU2], 125
Lycaena chrysaeis; [CoL]
Lycaena chryseis; [CoL]
Lycaena hippothoe euridice; [iNaturalist]
Lycaena hippothoe hippothoe; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [iNaturalist]
Lycaena subtusimpunctata; [CoL]
Polyommatus chrysaeis[sic, chryseis?]; de Selys, 1845, Mém. Soc. Roy. Sci. Liége 2: 3
Polyommatus chryseis confluens; Oberthür, 1896, Étud. d'Ent. 20: pl. 5, f. 80

L. h. stiberi (Gerhard, 1852) N.Scandinavia, N.Urals

Polyommatus chryseis var. stiberi Gerhard, 1852; Versuch Mon. europ. Schmett. (9): 19, pl. 35, f. 1a-b; TL: Lappland

Heodes hippothoe stiberi; [BRU2], 125; [BARU](note) ,
Lycaena hippothoe stiberi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Palaeochrysophanus hippothoe stiberi; [H&R], 249; [CoL]

L. h. eurydame (Hoffmannsegg, 1806) Alps?

Papilio eurydame Hoffmannsegg, 1806; Mag. f. Insektenk. 5: 178; TL: Mountains near Geneva

Lycaena hippothoe eurydame; [iNaturalist] ; [Heiner Ziegler] ; [Mario Maier]
Palaeochrysophanus hippothoe eurydame; [H&R], 249; [CoL]

L. h. eurybia (Ochsenheimer, 1808) Altai, Sayan, Far East

Papilio eurybia Ochsenheimer, 1808; Schmett. Europa 1 (2): 81
Chrysophanus hippothoe var. eurybia ab. wallentini Hirschke, 1910; Verh. zool.-bot. Wien 60: 412
sajana (Kozhanchikov, 1923) (Chrysophanus); Jb. Martjanov Staatmus. Minussinsk 1 (1): 10

Heodes hippothoe eurybia; [BRU2], 125
Heodes hippothoe sajana; [BARU](note) ,
Lycaena hippothoe sajana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [iNaturalist]

501x395 (~37Kb) ♂ ♀ underside An alpine meadow in the valley of the headwater of the Chikty Rivulet, 2500 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 22nd July 1998 (on Pedicularis anthemifolia Fisch. ex Colla), Photo © Oleg Kosterin

439x400 (~38Kb) ♂ underside A meadow in a depression on the Adon-Chelon elevation, at the mountain Tsagan-Obo, 10 km NNW of the village Tasyrkhoi, S Chita Province (Dahuria), Transbaikalia, Siberia, Russia. 9th July 1996, Photo © Oleg Kosterin

L. h. amurensis (Staudinger, 1892) Transbaikalia, Amur, Ussuri

Polyommatus hippothoe var. amurensis Staudinger, 1892; in Romanoff, Mém. Lépid. 6: 155

Heodes hippothoe amurensis; [BARU](note) , ; [BRU2], 125
Lycaena hippothoe amurensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [RFEL]
Palaeochrysophanus hippothoe amurensis; [MRS], 664; [CoL]

L. h. cisalpina (Fruhstorfer, 1909)

Chrysophanus hippothoe cisalpina Fruhstorfer, 1909; Int. ent. Zs. 3 (21): 120; TL: Tessin, Val Maggia and Fusio

L. h. engadiniana (Beuret, 1953)

Palaeochrysophanus hippothoe engadiniana Beuret, 1953; Mitt. ent. Ges. Basel (n.f.) 3 (1): (6-7)

Palaeochrysophanus hippothoe engadiniana; [CoL]

Lycaena candens (Herrich-Schäffer, [1844])

SEEU. See [maps]

Polyommatus candens Herrich-Schäffer, [1844]; Syst. Bearb. Schmett. Europ. 1 (7): (ii) f. 229-231; TL: Kleinasien [Turkey]
- = hippothoe; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 91
Polyommatus candens var. dido Gerhard, 1850; Versuch Mon. europ. Schmett. (2): 5, pl. 8, f. 2a-b
- = hippothoe; Bridges, 1994, Cat. Riodinidae & Lycaenidae: (VIII) 139

Heodes candens; [BRU2]: 125, pl. 60, f. 13-15
Lycaena (Heodes) candens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Lycaena candens; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE]
Palaeochrysophanus candens; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [CoL]

L. c. leonhardi (Fruhstorfer, 1917)

Chrysophanus hippothoe leonhardi Fruhstorfer, 1917; Ent. Rundschau 34 (4): 16; TL: Bulgaria, "Rilo..., Herzegowina Vucijabara"

Palaeochrysophanus hippothoe leonhardi; [H&R], 250; [CoL]
leonhardi; Abadjiev, 2002, Neue Ent. Nachr. 53: 20

L. c. pfeifferi (Beuret, 1952) Caucasus, Transcaucasia

Palaeochrysophanus candens pfeifferi Beuret, 1952; Mitt. ent. Ges. Basel (n.f.) 2 (7): 59; TL: Caucasus

Heodes candens pfeifferi; [BRU2], 125
Palaeochrysophanus candens pfeifferi; [CoL]

Subgenus Phoenicurusia Verity, 1943

Phoenicurusia Verity, 1943; Le Farfalle diurn. d'Italia 2: 21; TS: Polyommatus phoenicurus Lederer [= Polyommatus dimorphus var. margelanica Staudinger]
Margelycaena Kemal, 2006; Priamus Suppl. 2: 81 (repl. Phoenicurusia Verity, 1943); TS: Polyommatus dimorphus var. margelanica Staudinger
- = Phoenicurusia; Krupitsky, 2017, Zootaxa 4306 (1): 138

Subgenus Phoenicurusia (Lycaena); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Subgenus Phoenicurusia (Tharsalea); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Margelycaena; [iNaturalist] ; [NHM-BMW]
Phoenicurusia; Tuzov, 2016, Far Eastern Ent. 310: 11; [CoL] ; [iNaturalist] ; [NHM-BMW]
Phoenicurusia (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441
Phoenicurusia (Lycaenidae); [BRU2], 130
Phoenicurusia (Lycaenini); Krupitsky, 2017, Zootaxa 4306 (1): 138 (list)

Lycaena margelanica Staudinger, 1881

S.Tajikistan - N.Tien-Shan. See [maps]

Polyommatus dimorphus var. margelanica Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 282; TL: Margelan [northern slope of Alaisky Mts., Kirghizia]

Lycaena (Phoenicurusia) margelanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Lycaena margelanica; Nekrutenko & Effendi, 1983, Vestn. zool. 1983 (4): 12; Nekrutenko, 1984, Vestn. zool. 1984 (6): 44; [iNaturalist]
Margelycaena margelanica; Kemal, 2006, Priamus Suppl. 2: 81
Phoenicurusia margelanica; [BRU2]: 130, pl. 60, f. 16-24; [iNaturalist] ; [LEPIFORUM]
Polyommatus phoenicurus var. margelanica; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 361

L. m. margelanica Ghissar, Darvaz, Alai, W.Tien-Shan, W.Pamir

Lycaena margelanica margelanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76; [iNaturalist]
Phoenicurusia margelanica margelanica; [BRU2], 130
Phoenicurusia phoenicurus margelanica; [CoL]

L. m. legezini (Zhdanko, 2000) S.Ghissar, Babatag, Karatau, Teriklitau, Rangontau

Phoenicurusia margelanica legezini Zhdanko, 2000; Tethys 1: 214

Lycaena margelanica legezini; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Phoenicurusia margelanica legezini; [BRU2], 130

L. m. nigra (Zhdanko, 2000) Zailiisky Alatau, Lake Issyk-Kul (mountains around)

Phoenicurusia margelanica nigra Zhdanko, 2000; Tethys Ent. Res. 2: 214

Lycaena margelanica nigra; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 76
Phoenicurusia margelanica nigra; [BRU2], 131

L. m. gaudbunda Korb, 2012 Kyrgyzstan

Lycaena margelanica gaudbunda Korb, 2012; Eversmannia 29-30: 10; TL: Kyrgyzstan, Terskey Alatau, 1700m

L. m. goshkoi (Tuzov, 2016) Tadjikistan (Darvaz)

Phoenicurusia margelanica goshkoi Tuzov, 2016; Far Eastern Ent. 310: 12; TL: Tadjikistan, Darvaz, Hozratishoch Mts, Dashtidzhum loc., 1200m

Lycaena euphratica Eckweiler, 1989

Turkey. See [maps]

Lycaena (Phoenicurusia) euphratica Eckweiler, 1989; Nachr. ent. Ver. Apollo NF 10 (2): 83; TL: Turkey, Bingöl, 14-24km S. Genç, 1200-1400m, 30km S Bingöl

Lycaena euphratica; ; [iNaturalist] ; [LEPIFORUM]
Margelycaena euphratica; Kemal, 2006, Priamus Suppl. 2: 81 (note); [iNaturalist]

Subgenus Hyrcanana Bethune-Baker, 1914

Hyrcanana Bethune-Baker, 1914; Ent. Rec. J. Var. 26 (5): 135; TS: Polyommatus caspius Lederer
Sarthusia Verity, 1943; Le Farfalle diurn. d'Italia 2: 20; TS: Polyommatus sarthus Staudinger
- = Hyrcanana; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; Nekrutenko, 1983, Vestn. zool. 1983 (3): 8
- = Subgenus Hyrcanana; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)

Subgenus Hyrcanana (Tharsalea); Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 13 (list)
Hircanana[sic, Hyrcanana];
Hyrcanana; Nekrutenko, 1983, Vestn. zool. 1983 (3): 8; [CoL] ; [iNaturalist] ; [NHM-BMW]
Hyrcanana (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Hyrcanana (Lycaenidae); [BRU2], 135
Hyrcanana (Lycaeninae); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78
Hyrcanana (Lycaenini); Krupitsky, 2017, Zootaxa 4306 (1): 138 (list)
Sarthusia; [CoL] ; [NHM-BMW]
Sarthusia (Section Lycaena); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 441

Lycaena caspius (Lederer, 1869)

Kopet-Dagh, Ghissar?, N.Iran, Afghanistan?. See [maps]

Polyommatus caspius Lederer, 1869; Horae Soc. ent. ross. 6 (2): 76, pl. 4, f. 3; TL: Astrabad [Gorgan, N.Iran]
?Lycaena afghana Howarth & Povolný, 1976; Èas. morav. Mus. Brn. 61: 150
- = Hyrcanana sartha ophion; Nekrutenko, 1983, Vestn. zool. 1983 (3): 14
- = Hyrcanana caspius; [BRU2], 135 (?)

Hyrcanana caspia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78
Hyrcanana caspius; [BRU2]: 135, pl. 61, f. 30-32; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [CoL] ; [iNaturalist]
Lycaena caspius; [BOW]: pl. 206, f. 4; [LEPIFORUM]

L. c. caspius Kopet-Dagh

Hyrcanana caspia caspia; Nekrutenko, 1983, Vestn. zool. 1983 (3): 11
Hyrcanana caspius caspius; [BRU2], 135

Lycaena evansii (de Nicéville, 1902)

Kashmir, Chitral. See [maps]

Chrysophanus evansii de Nicéville, 1902; J. Bombay nat. Hist. Soc. 14 (2): 249, pl. FF, f. 11; TL: Drosh, Chitral, Western Himalayas

Hyrcanana caspius evansi; Bálint, 1992, Linn. Belg. 13 (8): 400
Hyrcanana evansi; Nekrutenko, 1983, Vestn. zool. 1983 (3): 15
Lycaena evansi; [EBW]
Lycaena evansii; [CoL]

Misidentified/misapplied:

Hyrcanana evansi (= evansi); Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name)

Lycaena transiens (Staudinger, 1886)

Polyommatus caspius var. transiens Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 201
- = Hyrcanana caspius; [BRU2], 135 (?)

Hyrcanana caspia transiens; Nekrutenko, 1983, Vestn. zool. 1983 (3): 11
Hyrcanana transiens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78
Polyommatus caspius var. transiens; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 360

L. t. transiens

Hyrcanana transiens transiens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78

Lycaena ophion Hemming, 1933

Kopet-Dagh, Ghissar-Darvaz. See [maps]

Lycaena ophion Hemming, 1933; Entomologist 66: 278 (repl. caudatus Staudinger, 1901); TL: [Karateginsky Mts., Tajikistan]
Chrysophanus sarthus var. caudatus Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 3: 73 (preocc.); TL: Buch.[ara] s.[eptentrionalis] or.[ientalis] (Karategin)

Hyrcanana ophion; [BRU2]: 135, pl. 61, f. 33-35, 42-44; [iNaturalist]

L. o. ophion Ghissar-Darvaz

Hyrcanana ophion ophion; [BRU2], 135
Hyrcanana sartha ophion; Nekrutenko, 1983, Vestn. zool. 1983 (3): 14

L. o. meridianalis (Shchetkin, 1963) Tadzhikistan

ophion meridianalis (Shchetkin, 1963) (Chrysophanus); Trudy Inst. Zool. Parasit., Stalinabad 24: 52; TL: Khozratishoh Mts.
- = Hyrcanana sartha ophion; Nekrutenko, 1983, Vestn. zool. 1983 (3): 14

Hyrcanana ophion meridianalis; [BRU2], 135
Hyrcanana transiens meridianalis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78

Lycaena sartha (Staudinger, 1886)

Ghissar-Darvaz, Pamirs-Alai, NE.Afghanistan. See [maps]

Polyommatus sarthus Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 202; TL: Südlichen Alai-Gebirgen wohl bei Kara-Kasuk

Hyrcanana sartha sartha; Nekrutenko, 1983, Vestn. zool. 1983 (3): 14
Hyrcanana sartha; [BRU2]: 135, pl. 61, f. 39-41; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78; [iNaturalist]
Hyrcanana sarthus; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109; [LEPIFORUM]
Lycaena sarthus; [BOW]: pl. 206, f. 4 (text)
Polyommatus sarthus; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 361, pl. 6, f. 5

L. s. hofmanni (ten Hagen, 2006) Iran

Hyrcanana sartha hofmanni ten Hagen, 2006; Nachr. Ent. Ver. Apollo NF 27 (3): 109; TL: Iran, Esfahan, Kuh-e Karkas, 2500m

Hyrcanana sartha hofmanni; Alipanah, 2017, SHILAP Revta lepid. 44 (174): 197

Lycaena pamira (Nekrutenko, 1983)

W.Pamirs. See [maps]

Hyrcanana pamira Nekrutenko, 1983; Vestn. zool. 1983 (3): 12, f. 4; TL: [Nyut, Ishakshimsky Mts., SW.Pamirs, Tajikistan]

Hyrcanana pamira; [BRU2]: 136, pl. 61, f. 45-47; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78

Lycaena sultan (Lang, 1884)

W.Ghissar (Zeravshansky, Ghissarsky, Baisuntau, Fanskie). See [maps]

Polyommatus sultan Lang, 1884; Rhopalocera Europa Descripta et Delineata 1: 368 (Stgr.); TL: Samarkand
Polyommatus sultan Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 202 (preocc. Polyommatus sultan Lang, 1884); TL: Samarkand
Chrysophanus sultani Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 75 (repl. Polyommatus sultan Staudinger, 1886); TL: Samarkand

Hyrcanana sultan; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name); Nekrutenko, 1983, Vestn. zool. 1983 (3): 11; [BRU2]: 136, pl. 61, f. 36-38; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 78
Lycaena sultan; [EBW]; [BOW]: pl. 206, f. 4 (text)
Polyommatus sultan; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 360, pl. 6, f. 6a-b; [CoL]

Lycaena svenhedini (Nordström, 1935)

S.Gansu, Shaanxi, N.Sichuan. See [maps]

Chrysophanus svenhedini Nordström, 1935; Ark. Zool. 27A (7): 30; TL: S.Kansu

Lycaena svenhedini; [CoL] ; [iNaturalist]

Lycaena irmae Bailey, 1932

S.Tibet. See [maps]

Lycaena irmae Bailey, 1932; J. Bombay nat. Hist. Soc. 35: 697; TL: Gyantse, Tibet, 13000ft

Lycaena irmae; [EBW], (name); [CoL] ; [iNaturalist]

Lycaena li (Oberthür, 1886)

W.China. See [maps]

Chrysophanus li Oberthür, 1886; Bull. Soc. ent. Fr. (6) 6: xxii; TL: Ta-tsien-lou
Chrysophanus li var. vernalis Oberthür, 1886; Bull. Soc. ent. Fr. (6) 6: xxiii
Chrysophanus li var. aestivalis Oberthür, 1886; Bull. Soc. ent. Fr. (6) 6: xxiii

Chrysophanus li; Oberthür, 1914, Étud. Lépid. Comp. 9 (2): 55, pl. 255, f. 2152
Helleia li; [MRS], 662; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109
Lycaena li; [EBW]; [BOW]: pl. 206, f. 5, 6; [CoL] ; [iNaturalist]
Lycaena vernalis; [CoL]

L. l. lilacina (Oberthür, 1914)

Chrysophanus li lilacina Oberthür, 1914; Étud. Lépid. Comp. 9 (2): 55, pl. 255, f. 2153

Lycaena li lilacina; [CoL]

Lycaena ouang (Oberthür, 1891)

Yunnan. See [maps]

Chrysophanus ouang Oberthür, 1891; Étud. d'Ent. 15: 17, pl. 2, f. 19; TL: Tsekou, NW. Yunnan

Helleia ouang; Sibatani, 1974, Aust. ent. Soc. 1974 (13): 109 (name)
Lycaena ouang; [EBW], (name); Huang, 2001, Neue Ent. Nachr. 51: 75 (note), pl. 3, f. 24; [CoL] ; [iNaturalist]

L. o. nujiangensis Huang, 2001

Lycaena ouang nujiangensis Huang, 2001; Neue Ent. Nachr. 51: 75, pl. 3, f. 20, pl. 4, f. 25; TL: Gamagou

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Goto:

1000x1272 (~143Kb) ♀ upperside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1000x1380 (~170Kb) ♀ underside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1000x1235 (~144Kb) ♀ USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela
1000x1216 (~133Kb) ♂ USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1000x1291 (~152Kb) ♂ USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1000x1153 (~137Kb) ♂ USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela
1400x1333 (~228Kb) ♂ ♀ USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela

700x885 (~45Kb) underside USA: Tolmie State Park (47°07'23N 122°46'39W), Thurston Co., WA, 12.7.1999, Photo © Markku Savela	700x597 (~66Kb) underside USA: nr Crystal Mountain / FR9726 (47°13'56N 120°38'25W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela	600x785 (~68Kb) USA: Teanaway, Jungle Creek, (47°20'06N 120°55'21W ±0.5km) Kittitas Co., WA, 21.7.2000, Photo © Markku Savela
500x546 (~35Kb) ♂ USA: FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°19'58N 121°42'43W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela	900x1059 (~144Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W ~ ±0.5km), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela	900x1241 (~147Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W ~ ±0.5km), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

700x951 (~56Kb) USA: Table Mtn, (~ 47°19N 120°42W), Kittitas Co., WA, 30.7.1999, Photo © Markku Savela	660x623 (~60Kb) underside USA: FR9705, Table Mtn (47°18'20N 129°41'43W), Kittitas Co., WA, 19.7.1999, Photo © Markku Savela	900x728 (~61Kb) ♂ upperside USA: Teanaway, Jungle Creek, (47°20'06N 120°55'21W ±0.5km) Kittitas Co., WA, 21.7.2000, Photo © Markku Savela
600x717 (~58Kb) underside USA: Teanaway, Jungle Creek, (47°20'06N 120°55'21W ±0.5km) Kittitas Co., WA, 21.7.2000, Photo © Markku Savela	1000x1196 (~119Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W ~ ±0.5km), Kittitas Co., WA, 29.7.2006 (on Sedum divergens), Photo © Markku Savela

462x324 (~64Kb) ♀ upperside Awarua Bay, nr Invercargill, Southland. South Island, Jan 2006, Photo © Christopher A. Rickards	483x324 (~51Kb) ♀ underside Awarua Bay, nr Invercargill, Southland. South Island, Jan 2006, Photo © Christopher A. Rickards	478x324 (~63Kb) ♂ upperside Awarua Bay, nr Invercargill, Southland. South Island, Jan 2006, Photo © Christopher A. Rickards
457x324 (~45Kb) ♂ underside Awarua Bay, nr Invercargill, Southland. South Island, Jan 2006, Photo © Christopher A. Rickards

489x324 (~81Kb) ♀ upperside Milford Sound, Fiordland, South Island, NZ, Jan 2006, Photo © Christopher A. Rickards	482x324 (~68Kb) ♀ underside Milford Sound, Fiordland, South Island, NZ, Feb 2006, Photo © Christopher A. Rickards	458x324 (~77Kb) ♂ upperside Milford Sound, Fiordland, South Island, NZ, Jan 2006, Photo © Christopher A. Rickards
498x324 (~57Kb) ♂ underside Milford Sound, Fiordland, South Island, NZ, Feb 2006, Photo © Christopher A. Rickards

375x564 (~64Kb) ♀ upperside New Zealand, Malborough Sounds region, South Island, 13 Nov 2005, Photo © Christopher A. Rickards	564x375 (~56Kb) ♂ upperside New Zealand, Malborough Sounds region, South Island, 13 Nov 2005, Photo © Christopher A. Rickards	564x375 (~49Kb) ♂ underside New Zealand, Malborough Sounds region, South Island, 13 Nov 2005, Photo © Christopher A. Rickards
462x324 (~52Kb) ♂ Homer Tunnel, Fiordland, South Island, NZ, Feb 2006, Photo © Christopher A. Rickards	487x324 (~36Kb) ♀ underside Catlins, South Island, NZ, Feb 2006, Photo © Christopher A. Rickards	324x466 (~25Kb) ♂ underside Dunedin, South Island, NZ, Feb 2007, Photo © Christopher A. Rickards
487x324 (~41Kb) ♀ upperside Banks Penninsular, South Island, NZ, Dec 2005, Photo © Christopher A. Rickards