Life Insecta Lepidoptera Ditrysia Papilionoidea Lycaenidae Polyommatinae

600x400 (~104Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~101Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~79Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar
600x400 (~49Kb) India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

• Acytolepis puspa (Horsfield, [1828]) Peninsular India, Himalayas, Assam, Ceylon, Burma, Philippines, Nicobars, New Guinea. See [maps] Common Hedge Blue

  • Polyommatus puspa Horsfield, [1828]; Descr. Cat. lep. Ins. Mus. East India Coy (1): 67; TL: Java
  • Cyaniris puspa var. watasei Matsumura, 1915; Ent. Mag. Kyoto 1 (2): 66, 68

  • Acytolepis puspa prominans; [CoL]
  • Acytolepis puspa; [MRS], 679; [BMP]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161; [CoL] ; [iNaturalist] ; [LEPIFORUM]
  • Celastrina puspa; [BOW]: pl. 175, f. 20, 21
  • Cyaniris puspa; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 572; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 284
  • Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 437; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 24; [BIR], 274

  600x400 (~59Kb) underside India: Subansiri, 31.10.2005, Photo © Kedar Tokekar

  600x400 (~75Kb) underside India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

  Larva on Glochidion fortunei, Rhododendron spp., Distylium racemosum, Rosa multiflora, R. wichuraiana, R. centifolia, Prunus zippeliana, Glochidion obovatum, Myrica rubra, Quercus phillyreoides, Celtis sinensis, Astilbe thunbergii [MRS], Schleichera oleosa, Hiptage benghalensis, Xylia dolabriformis [BMP], Viburnum , Combretaceae , Cycadaceae , Shorea , Enkianthus , Lyonia , Rhododendron , Vaccinium , Antidesma , Breynia , Bridelia , Glochidion , Calliandra , Cylista , Paracalyx , Pellophorum , Xylia , Quercus , Distylum , Hiptage , Myrica , Aruncus , Chaenomeles , Malus , Photinia , Prunus , Rosa , Spiraea , Dimocarpus , Euphoria , Litchi , Sapindus , Schleichera , Astilbe , Celtis Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

  • A. p. puspa

    • Lycaenopsis puspa puspa; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. cagaya (C. & R. Felder, [1865]) Philippines (Luzon)

    • Lycaena cagaya C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 278, pl. 34, f. 11-13
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Cyaniris puspa ottonis Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287; TL: Palawan

    • Acytolepis puspa cagaya; [CoL] ; [iNaturalist]
    • Acytolepis puspa ottonis; [CoL]
    • Lycaenopsis puspa cagaya; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Lycaenopsis puspa ottonis; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. lambi (Distant, 1882) Siam, Vietnam, Tenasserim

    • Polyommatus lambi Distant, 1882; Ann. Mag. nat. Hist. (5) 10 (57): 245; TL: Malay Peninsula
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Cyaniris imperatrix Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 444; TL: Thailand
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa imperatrix; [CoL]
    • Acytolepis puspa lambi; [BMP]: 240, pl. 35, f. 44-46; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
    • Lycaenopsis puspa imperatrix; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Lycaenopsis puspa lambi; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. puspinus (Kheil, 1884) Nias

    • puspa puspinus (Kheil, 1884) (Plebeius); Rhopalocera Ins. Nias: 30, f. 37-39; TL: Nias
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa puspinus; [CoL]
    • Cyaniris puspinus; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287
    • Lycaenopsis puspa puspinus; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. kuehni (Röber, 1886) Sulawesi

    • Plebeius kühni Röber, 1886; Corr.-Bl. Ent. Ver. Iris 1 (3): 60, pl. 4, f. 29 (partim); TL: E.Sulawesi
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Lycaenopsis puspa martini Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 870; TL: Sulawesi
      • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Lycaenopsis puspa hyllus Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 870; TL: Banggai
      • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Cyanirus[sic] kühni kalawari Ribbe, 1926; Ent. Mitt. 15 (1): 89; TL: Sulawesi, Kalawara
      • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Cyaniris kühni bangkaiensis Ribbe, 1926; Ent. Mitt. 15 (1): 89; TL: Banggai
      • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

    • Acytolepis puspa hyllus; [CoL]
    • Acytolepis puspa kuehni; Toxopeus, 1928, Tijdschr. Ent. 71: 231; Eliot & Kawazoé, 1983, Butts. Lycaenopsis group: 180; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
    • Acytolepis puspa kühni; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54, f. 23-24
    • Acytolepis puspa martini; [CoL]
    • Cyaniris kuhni kalawari; [CoL]
    • Cyaniris kuhni; [CoL]
    • Cyaniris kuhni[sic, kuehni]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287
    • Lycaenopsis puspa kuehni; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Lycaenopsis puspa kühni; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. cyanescens (de Nicéville, 1890) Nicobar Is.

    • Cyaniris cyanescens de Nicéville, 1890; Butts India Burmah Ceylon 3: 103
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa cyanescens; [CoL]
    • Lycaenopsis puspa cyanescens; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. prominens (de Nicéville, 1890)

    • Cyaniris cyanescens prominens de Nicéville, 1890; Butts India Burmah Ceylon 3: 103

    • Lycaenopsis puspa prominens; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. dammae (Heron, 1894) Timor, Damar, Wetar

    • Cyaniris dammae Heron, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 104; TL: Damma I.
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa dammae; [BAUR], 383; [CoL]
    • Cyaniris puspa damae[sic, dammae]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 286
    • Lycaenopsis puspa dammae; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. timorensis (Butler, 1900) Timor

    • Cyaniris timorensis Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 443; TL: Timor
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa timorensis; [CoL]
    • Cyaniris puspa timorensis; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 286 (note)
    • Lycaenopsis puspa timorensis; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. splendens (Butler, 1900)

    • Cyaniris splendens Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 444; TL: Larut Hills, Perak
      • = Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa splendens; [CoL]
  • A. p. myla (Fruhstorfer, 1909) Formosa

    • Cyaniris puspa myla Fruhstorfer, 1909; Ent. Zs. 23 (25): 116; TL: Formosa
    • Cyaniris postimacula Matsumura, 1929; Insecta Matsumurana 3 (2/3): 104; TL: Formosa
    • myla (Matsumura, 1929) (Cyaniris); Insecta Matsumurana 3 (2/3): 104 (preocc. Fruhstorfer)
    • Celastrina puspa formosana Bethune-Baker, 1914; Ent. Mitt. 3 (4): 127; TL: Formosa

    • Acytolepis puspa myla; [MRS], 679; [CoL] ; [iNaturalist]
    • Cyaniris puspa myla; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 284
    • Lycaenopsis puspa myla; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 24

    Larva on Glochidion acuminatum, G. lanceolatum, G. rubrum, Bridelia balansae, Euphoria longan, Sapindus mukorossi, Vaccinium oldhamii, V. smallii, Enkyathe perulatus, Lyonia ovalifolia, Viburnum dilatatum, Prunus padus, Chaenomeles speciosa, Malus domestica, Spiraea cantoniensis, Aruncus dioicus, Astilbe thunbergii [MRS]

  • A. p. hermagoras (Fruhstorfer, 1910) Hainan

    • Cyaniris puspa hermagoras Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 284; TL: Hainan

    • Acytolepis puspa hermagoras; [MRS], 679; [CoL]
    • Lycaenopsis puspa hermagoras; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. sania (Fruhstorfer, 1910)

    • Cyaniris puspa sania Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285

    • Acytolepis puspa sania; [CoL]
    • Lycaenopsis puspa sania; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. gisca (Fruhstorfer, 1910) NE.Afghanistan - Burma, Thailand, Cambodia, Laos, Vietnam, S.China, Yunnan

    • Cyaniris puspa gisca Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285; TL: Sikkim
    • Lycaenopsis puspa gisca f. artena Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25; TL: India
    • Lycaenopsis puspa telis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25; TL: Andaman
    • Celastrina puspa barneyi Corbet, 1941; Hongkong Nat. 10 (3/4): 149; TL: Hong Kong

    • Acytolepis puspa artena; [CoL]
    • Acytolepis puspa barneyi; [CoL]
    • Acytolepis puspa gisca; [MRS], 679; Huang, 2003, Neue Ent. Nachr. 55: 106 (note); [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
    • Acytolepis puspa telis; [CoL]
    • Lycaenopsis puspa gisca; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25

    848x752 (~135Kb) ♂ underside Cambodia, Ratanakiri Province, Virachey National Park biffer zone, Yorn village environs, Yak Kae Rivulet, 4th June 2013, Photo © Oleg Kosterin

    813x965 (~262Kb) underside Cambodia, Mondukliri Province, the Pulung (?) river 12 km NE of Sen Monorom, between Srae Empum and Buu Sraa, 25th June 2018, Photo © Oleg Kosterin

  • A. p. hermione (Fruhstorfer, 1905) Bawean

    • Cyaniris puspa hermione Fruhstorfer, 1905; Soc. Ent. 20 (1): 2; TL: Bawean

    • Acytolepis puspa hermione; [CoL]
    • Cyaniris puspa hermione; Fruhstorfer, 1905, Ent. Zs. 19 (12): 64, f. 10
    • Lycaenopsis puspa hermione; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. contilia (Fruhstorfer, 1910) Sumbawa

    • Cyaniris puspa contilia Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285; TL: Sumbawa, Sumba

    • Acytolepis puspa contilia; [CoL]
    • Lycaenopsis puspa contilia; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. bazilana (Fruhstorfer, 1910) Philippines (Mindoro, ...?)

    • Cyaniris puspa bazilana Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Bazilan
    • Cyaniris puspa georgi Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Camiguin de Mindanao
    • Lycaenopsis puspa sabis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25

    • Acytolepis puspa bazilana; [iNaturalist]
    • Acytolepis puspa georgi; [CoL]
    • Acytolepis puspa sabis; [CoL]
    • Lycaenopsis puspa bazilana; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Lycaenopsis puspa georgi; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. calata (Fruhstorfer, 1910) Flores

    • Cyaniris puspa calata Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Flores

    • Acytolepis puspa calata; [CoL]
    • Lycaenopsis puspa calata; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • ?Udara placidula calata;
  • A. p. darmis (Fruhstorfer, 1917) Kai, Aru?

    • Lycaenopsis puspa darmis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 27; TL: Key I.

    • Acytolepis puspa darmis; [BAUR], 383 (text)
  • A. p. mygdonia (Fruhstorfer, 1917) Sumatra, Borneo

    • Lycaenopsis puspa mygdonia Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 26; TL: N.Sumatra; Borneo

    • Acytolepis puspa mygdonia; [CoL]
  • A. p. roberi (Fruhstorfer, 1917) Goram

    • Lycaenopsis puspa röberi Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 27; TL: Goram

    • Acytolepis puspa roberi; [BAUR], 383 (text); [CoL]
  • A. p. volumina (Fruhstorfer, 1922)

    • Lycaenopsis volumina Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 869, Lfg. (302)

    • Acytolepis puspa volumina; [CoL]
  • A. p. tymbria (Fruhstorfer, 1922)

    • puspa tymbria (Fruhstorfer, 1922) (Lycaenopsis); in Seitz, Gross-Schmett. Erde 9: 870

    • Acytolepis puspa tymbria; [CoL]
  • A. p. deronda (Fruhstorfer, 1922) Sula Is.

    • Lycaenopsis puspa deronda Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 871; TL: Sula Is
    • Lycaenopsis puspa var. Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa deronda; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162; [CoL]
  • A. p. vandeldeni Toxopeus, 1929 Sumatra

    • Acytolepis puspa vandeldeni Toxopeus, 1929; Tijdschr. Ent. 72 (3/4): 207; TL: Pulu Weh

    • Acytolepis puspa vandeldeni; [CoL]
  • A. p. felderi Toxopeus, 1927 Ceylon

    • Acytolepis puspa felderi Toxopeus, 1927; Tijdschr. Ent. 70 (3/4): 294

    • Acytolepis puspa felderi; [CoL]
  • A. p. ishigakiana (Matsumura, 1929)

    • Cyaniris ishigakiana Matsumura, 1929; Insecta Matsumurana 3 (2/3): 104; TL: Okinawa

    • Acytolepis puspa ishigakiana; [iNaturalist]
    • Celastrina ishigakiana; [CoL]
  • A. p. umenonis (Matsumura, 1931) Japan

    • Lycaenopsis (Cyaniris) puspa f. umenonis Matsumura, 1931; Insecta Matsumurana 6 (1-2): 44

    • Acytolepis puspa umenonis; [CoL]
    • Celastrina puspa umenonis; Kudrna, 1974, Atalanta 5: 115
  • A. p. haani (Kalis, 1938) Kangean I.

    • puspa haani (Kalis, 1938) (Lycaenopsis); Ent. Rdsch. 55: 198; TL: Kangean I.

    • Acytolepis puspa haani; [CoL]
  • A. p. siporana (Riley, [1945]) Mentawi Is.

    • puspa siporana (Riley, [1945]) (Celastrina); Trans. R. ent. Soc. Lond. 94 (2): 256

    • Acytolepis puspa siporana; [CoL]
• Acytolepis armenta (Fruhstorfer, 1910) Borneo. See [maps]

  • Cyaniris armenta Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287; TL: Borneo; Lawas

  • Cyaniris armenta; [CoL]
• Acytolepis lilacea (Hampson, 1889) Nilgiris, Palnis, Coorg. See [maps] Hampson's Hedge Blue

  • Cyaniris puspa var. lilacea Hampson, 1889; J. asiat. Soc. Bengal 57 (4): 356; TL: Nilgiris and Nellyampathy Hills, Cochin

  • Acytolepis lilacea; [iNaturalist]
  • Acytolepis puspa lilacea; [CoL]
  • Lycaenopsis lilacea; [BIR], 274
  • A. l. lilacea

    • Cyaniris puspa lilacina[sic, lilacea?]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 285
    • Lycaenopsis puspa lilacea; ; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. l. moorei (Toxopeus, 1926) Ceylon

    • Lycaenopsis moorei Toxopeus, 1926; Treubia 8 (3-4): 365

    • Acytolepis puspa moorei; [CoL]
  • A. l. indochinensis Eliot & Kawazoé, 1983 Burma, Thailan, Laos

    • Acytolepis lilacea indochinensis Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 187, f. 135, 268, 284; TL: Vang Vien, Laos

    • Acytolepis lilacea indochinensis; [Yutaka Inayoshi]
• Acytolepis ripte (Druce, 1895) Borneo. See [maps]

  • Cyaniris ripte Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 574, pl. 32, f. 11; TL: Labuan

  • Acytolepis ripte; [iNaturalist] ; [LEPIFORUM]
  • Cyaniris ripte; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 301; [CoL]
  • Lycaenopsis rifte; [CoL]
  • Lycaenopsis ripte; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 434; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 33
• Acytolepis najara (Fruhstorfer, 1910) Sulawesi. See [maps]

  • Cyaniris najara Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287 (part, ♂); TL: Sulawesi, Bua-Krang, 5000ft

  • Acytolepis najara; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162; [iNaturalist]
  • Cyaniris najara; [CoL]

  Misidentified/misapplied:

  • Cyaniris najara (= Acytolepis samanga); Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287 (part, ♀)
• Acytolepis samanga (Fruhstorfer, 1910) Samanga

  • Cyaniris samanga Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Samangi, Bua
  • Lycaenopsis akasa ssp. Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 9
  • Lycaenopsis akasa parakasa Fruhstorfer, 1922; Gross-Schmett. Erde 9: 862; TL: Sulawesi, Minahassa
    • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55

  • Acytolepis puspa samanga; Toxopeus, 1928, Tijdschr. Ent. 71: 231 (partim, female); [CoL]
  • Acytolepis samanga; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162; [iNaturalist]
  • Celastrina samanga; Eliot, 1956, Bull. Raffles Mus. 27: 35, f. 2

24.5.2023 (6)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.